Mesosa

Subfamily

Lamiinae

Diagnosis - adult

Body length: 8–17 mm. 
Eyes: eye interommatidial setaeseta:
a sclerotized hair-like projection of the cuticle
absent, eye deeply emarginateemarginate:
notched at the margin
> half width or divided-max one row of facets between, eye ommatidial density fine. 
Antennaeantenna:
in larval and adult insects, paired segmented appendages, borne one on each side of the head, functioning as sense organs and bearing a large number of sensilla
: antennal length reaches between basebase:
the part of any appendage or structure that is nearest the body
and end of elytraelytron:
the leathery forewing of beetles, serving as a covering for the hind wings, commonly meeting opposite elytron in a straight line down the middle of the dorsum in repose
or reaching/surpassing end of body, antennal flagellar segments elongateelongate:
much longer than wide
, scapescape:
the first proximal segment of the antenna
cicatrix (scar) at apexapex:
end of any structure distad to the base
, antennal segment 3 > scapescape:
the first proximal segment of the antenna

Pronotumpronotum:
the upper and dorsal part of the prothorax
: pronotumpronotum:
the upper and dorsal part of the prothorax
shape transversetransverse:
broader than long
, pronotumpronotum:
the upper and dorsal part of the prothorax
lateral armature absent or blunt tubercule. 
Prosternum: prosternal processprosternal process:
a posterior extension of the prosternum between the coxae
dilated at apexapex:
end of any structure distad to the base
, procoxal cavities closed posteriorly. 
Elytraelytron:
the leathery forewing of beetles, serving as a covering for the hind wings, commonly meeting opposite elytron in a straight line down the middle of the dorsum in repose
: elytral length reaching or close to end of abdomen, elytral apicesapex:
end of any structure distad to the base
rounded or truncatetruncate:
cut off squarely at the tip
, elytral color pattern present. 
Legs: visible tarsomerestarsomere:
subdivision or article of the tarsus, usually numbering from two to five
: 4, femora clavateclavate:
thickening gradually toward the tip
, rarely robust, protibial spursprotibial spur:
sclerotized spine(s) located at the distal tibia; can be single, double, or absent
: 2, tarsal clawstarsal claw:
usually paired claws of the pretarsus, at the distal end of the leg
simple. 

Characterized by comparatively short, broad body. Head short, not narrower than pronotumpronotum:
the upper and dorsal part of the prothorax
, with produced antennal tubercleantennal tubercle:
the protuberance on the head where the antenna arises
, frontally with broad median longitudinal groove. Eyes highly emarginateemarginate:
notched at the margin
, divided almost into two equal lobes, lower lobelobe:
a rounded projection or protuberence
slightly larger than upper. Antennaeantenna:
in larval and adult insects, paired segmented appendages, borne one on each side of the head, functioning as sense organs and bearing a large number of sensilla
markedly longer than body (male) or barely extending beyond apexapex:
end of any structure distad to the base
of elytraelytron:
the leathery forewing of beetles, serving as a covering for the hind wings, commonly meeting opposite elytron in a straight line down the middle of the dorsum in repose
, on inner lower side with long dense bristles. Pronotumpronotum:
the upper and dorsal part of the prothorax
transversetransverse:
broader than long
, parallel-sided, narrower than elytraelytron:
the leathery forewing of beetles, serving as a covering for the hind wings, commonly meeting opposite elytron in a straight line down the middle of the dorsum in repose
, basally and apically without sharp flange. Elytraelytron:
the leathery forewing of beetles, serving as a covering for the hind wings, commonly meeting opposite elytron in a straight line down the middle of the dorsum in repose
short, broad, apically jointly rounded, in anterior third with coarse, simple or granular (M. curculionoides (L.)) punctationpunctation:
pits or depression of variable size in cuticle
. Legs not long, almost uniformly developed, femora sufficiently thickened. Sternite V uniformly, not highly, convex, with median longitudinal troughlike groove (female) or without it (male) (Cherepanov 1990Cherepanov 1990:
Cherepanov AI. 1990. Cerambycidae of Northern Asia, Vol. 3. Lamiinae, Part I. Oxonian Press, New Delhi 4: i–xiii + 1–300, 120 figs.
).

Large and robust insects. Antennaeantenna:
in larval and adult insects, paired segmented appendages, borne one on each side of the head, functioning as sense organs and bearing a large number of sensilla
largely exceeding the apexapex:
end of any structure distad to the base
of the elytraelytron:
the leathery forewing of beetles, serving as a covering for the hind wings, commonly meeting opposite elytron in a straight line down the middle of the dorsum in repose
in the male, reaching it in the female; article 1 with a sharp, oblique rim on the outer side of the apexapex:
end of any structure distad to the base
; article III longer than IV. Eyes very strongly indented. Pronotumpronotum:
the upper and dorsal part of the prothorax
transversetransverse:
broader than long
, weakly narrowed in front, not bearing a lateral spinespine:
a protuberance with an acute (sharp) distal end
. Elytraelytron:
the leathery forewing of beetles, serving as a covering for the hind wings, commonly meeting opposite elytron in a straight line down the middle of the dorsum in repose
rounded at the apexapex:
end of any structure distad to the base
(Villiers 1946Villiers 1946:
Villiers A. 1946. Coléoptères Cérambycides de l'Afrique du Nord. Faune de l'Empire Français, ORSC Paris 5: 1–152, 275 figs. http://www.cerambycoidea.com/titles/villiers1946.pdf
). Triangular shaped scapescape:
the first proximal segment of the antenna
.

Similar genera

Coptops, Mnemia, Japanomesosa, Eurymesosa, many Mesosini

Differential diagnosis

The enlarged apical scapescape:
the first proximal segment of the antenna
of this genus (and tribe) gives it a triangular shape that is unique. The squat body form of the tribe is also consistent. Within Mesosini, Mesosa has no spines on the antennomeresantennomere:
a subunit of the antenna, including the scape, pedicel, and flagellomeres
, the third antennomereantennomere:
a subunit of the antenna, including the scape, pedicel, and flagellomeres
obviously longer than the scapescape:
the first proximal segment of the antenna
, unarmed elytral apicesapex:
end of any structure distad to the base
, and the eyes are almost completely divided.

Distribution

Palearctic, Indomalaya

Recorded host plants

broadleaf; Abies, Larix, Pinus, Tsuga; Chamaecyparis

Remarks

7 spp. (Mesosa), 30 spp. (Aplocnemia), 2 spp. (Lissomesosa), 4 spp. (Metamesosa), 18 spp. (Perimesosa), 32 spp. (Saimia). Conifers: M. (Mesosa) japonicaM. (M.) mediofasciata, M. (Lissomesosa) kirisimana (Pinus), Mesosa (Metamesosa) nomurai (Abies, Pinus), Mesosa (Perimesosa) hirsuta (Abies, Pinus, broadleaf).

Synonyms

Dendrobius, Gistel, 1834 [for Mesosa]

Lamia (Mesosa) Brullé, 1832 [for Mesosa]

Haplocnemia Agassiz, 1846 [for Aplocnemia]

Helixoea Pascoe, 1865 [for Aplocnemia]

Taxonomy

Subgenus Mesosa, Latreille, 1829

Subgenus Aplocnemia, Stephens, 1831

Subgenus Lissomesosa, Yamasako and Ohbayashi, 2011

Subgenus Metamesosa, Breuning, 1938

Subgenus Perimesosa, Breuning, 1939

Subgenus Saimia, Pascoe, 1866

  Mesosa japonica  dorsal

Mesosa japonica dorsal

  Mesosa japonicanbsp; ventral

Mesosa japonica ventral

  Mesosa japonicanbsp; frontal

Mesosa japonica frontal

  Mesosa japonicanbsp; lateral

Mesosa japonica lateral

 PSG000960nbsp; Mesosa japonica  voucher
PSG000960 Mesosa japonica voucher