Sirex obesus

Taxonomy

Family: Siricidae
Subfamily: Siricinae
Genus: Sirex Linnaeus, 1760
Species: Sirex obesus Bradley, 1913
Common names: none

Background

Sirex obesus is a somewhat rare species from the southwestern United States with a stout, dark body (Schiff et al. 2012Schiff et al. 2012:
Schiff NM, Goulet H, Smith DR, Boudreault C, Wilson AD, and Scheffler BE. 2012. Siricidae (Hymenoptera: Symphyta: Siricoidea) of the Western Hemisphere. Canadian Journal of Arthropod Identification 21: 1-305.
).

Diagnostic characteristics

See Sirex for genus-level diagnostic characteristics.

Females:

  • hind femurfemur:
    the third segment of the leg between the trochanter and the tibia
    black (Schiff et al. 2012Schiff et al. 2012:
    Schiff NM, Goulet H, Smith DR, Boudreault C, Wilson AD, and Scheffler BE. 2012. Siricidae (Hymenoptera: Symphyta: Siricoidea) of the Western Hemisphere. Canadian Journal of Arthropod Identification 21: 1-305.
    )
  • abdomen abdomen:
    the third and last segment of an insect's body; in sawflies this is usually made up of 11 segments (segments 9 and 10 often fused)
    entirely black with metallic reflections (Schiff et al. 2012Schiff et al. 2012:
    Schiff NM, Goulet H, Smith DR, Boudreault C, Wilson AD, and Scheffler BE. 2012. Siricidae (Hymenoptera: Symphyta: Siricoidea) of the Western Hemisphere. Canadian Journal of Arthropod Identification 21: 1-305.
    )
  • all tibiaetibia:
    the fourth segment of the leg, between the femur and the tarsus
    and tarsitarsus:
    the fifth and last segment of the leg
    black (Schiff et al. 2012Schiff et al. 2012:
    Schiff NM, Goulet H, Smith DR, Boudreault C, Wilson AD, and Scheffler BE. 2012. Siricidae (Hymenoptera: Symphyta: Siricoidea) of the Western Hemisphere. Canadian Journal of Arthropod Identification 21: 1-305.
    )
  • fore wing fore wing:
    the anterior wing of each pair of wings; usually the largest wing of the pair
    tinted dark brown (Schiff et al. 2012Schiff et al. 2012:
    Schiff NM, Goulet H, Smith DR, Boudreault C, Wilson AD, and Scheffler BE. 2012. Siricidae (Hymenoptera: Symphyta: Siricoidea) of the Western Hemisphere. Canadian Journal of Arthropod Identification 21: 1-305.
    )
  • fore wing fore wing:
    the anterior wing of each pair of wings; usually the largest wing of the pair
    vein 3A absent (Schiff et al. 2012Schiff et al. 2012:
    Schiff NM, Goulet H, Smith DR, Boudreault C, Wilson AD, and Scheffler BE. 2012. Siricidae (Hymenoptera: Symphyta: Siricoidea) of the Western Hemisphere. Canadian Journal of Arthropod Identification 21: 1-305.
    )
  • gena gena:
    the area of the head between the compound eye and clypeus; also called the cheek
    pits large and dense, about 0–0.5 pit diameters apart (Schiff et al. 2012Schiff et al. 2012:
    Schiff NM, Goulet H, Smith DR, Boudreault C, Wilson AD, and Scheffler BE. 2012. Siricidae (Hymenoptera: Symphyta: Siricoidea) of the Western Hemisphere. Canadian Journal of Arthropod Identification 21: 1-305.
    )
  • vertex vertex:
    the dorsal portion of the head between the compound eyes, between the occiput and frons
    pits dense, about 0.3–0.4 times diameter of laterallateral:
    of or towards the side of the body
    ocellus (Schiff et al. 2012Schiff et al. 2012:
    Schiff NM, Goulet H, Smith DR, Boudreault C, Wilson AD, and Scheffler BE. 2012. Siricidae (Hymenoptera: Symphyta: Siricoidea) of the Western Hemisphere. Canadian Journal of Arthropod Identification 21: 1-305.
    )
  • second hind tarsomeretarsomere:
    a segment of the tarsus
    1.5 times as long as wide (Schiff et al. 2012Schiff et al. 2012:
    Schiff NM, Goulet H, Smith DR, Boudreault C, Wilson AD, and Scheffler BE. 2012. Siricidae (Hymenoptera: Symphyta: Siricoidea) of the Western Hemisphere. Canadian Journal of Arthropod Identification 21: 1-305.
    )
  • mesoscutum mesoscutum:
    the central and dorsal portion of the thorax between the scutellum and postnotum
    pits large, forming a few transverse ridges (Schiff et al. 2012Schiff et al. 2012:
    Schiff NM, Goulet H, Smith DR, Boudreault C, Wilson AD, and Scheffler BE. 2012. Siricidae (Hymenoptera: Symphyta: Siricoidea) of the Western Hemisphere. Canadian Journal of Arthropod Identification 21: 1-305.
    )
  • second hind tarsomeretarsomere:
    a segment of the tarsus
    with pulvilluspulvillus:
    soft pads used for surface adhesion, located in sawflies on the first 4 segments of the tarsus
    about 0.5–0.7 times length of tarsomeretarsomere:
    a segment of the tarsus
    (Schiff et al. 2012Schiff et al. 2012:
    Schiff NM, Goulet H, Smith DR, Boudreault C, Wilson AD, and Scheffler BE. 2012. Siricidae (Hymenoptera: Symphyta: Siricoidea) of the Western Hemisphere. Canadian Journal of Arthropod Identification 21: 1-305.
    )

Males:

  • hind femurfemur:
    the third segment of the leg between the trochanter and the tibia
    mostly reddish-brown (Schiff et al. 2012Schiff et al. 2012:
    Schiff NM, Goulet H, Smith DR, Boudreault C, Wilson AD, and Scheffler BE. 2012. Siricidae (Hymenoptera: Symphyta: Siricoidea) of the Western Hemisphere. Canadian Journal of Arthropod Identification 21: 1-305.
    )
  • gena gena:
    the area of the head between the compound eye and clypeus; also called the cheek
    completely blue-black with metallic reflections (Schiff et al. 2012Schiff et al. 2012:
    Schiff NM, Goulet H, Smith DR, Boudreault C, Wilson AD, and Scheffler BE. 2012. Siricidae (Hymenoptera: Symphyta: Siricoidea) of the Western Hemisphere. Canadian Journal of Arthropod Identification 21: 1-305.
    )
  • hind tibiatibia:
    the fourth segment of the leg, between the femur and the tarsus
    reddish-brown with dark band at the basebase:
    the beginning or most proximal area of any structure
    (Schiff et al. 2012Schiff et al. 2012:
    Schiff NM, Goulet H, Smith DR, Boudreault C, Wilson AD, and Scheffler BE. 2012. Siricidae (Hymenoptera: Symphyta: Siricoidea) of the Western Hemisphere. Canadian Journal of Arthropod Identification 21: 1-305.
    )
  • gena gena:
    the area of the head between the compound eye and clypeus; also called the cheek
    pits dense, about 1 diameter apart (Schiff et al. 2012Schiff et al. 2012:
    Schiff NM, Goulet H, Smith DR, Boudreault C, Wilson AD, and Scheffler BE. 2012. Siricidae (Hymenoptera: Symphyta: Siricoidea) of the Western Hemisphere. Canadian Journal of Arthropod Identification 21: 1-305.
    )
  • fore wing fore wing:
    the anterior wing of each pair of wings; usually the largest wing of the pair
    tinted yellow (Schiff et al. 2012Schiff et al. 2012:
    Schiff NM, Goulet H, Smith DR, Boudreault C, Wilson AD, and Scheffler BE. 2012. Siricidae (Hymenoptera: Symphyta: Siricoidea) of the Western Hemisphere. Canadian Journal of Arthropod Identification 21: 1-305.
    )
  • antennae completely black or black with brown pedicelpedicel:
    the second antennal segment, between the scape and flagellum
    and flagellomereflagellomere:
    A segment of the flagellum of the antenna
    1 and 2 (Schiff et al. 2012Schiff et al. 2012:
    Schiff NM, Goulet H, Smith DR, Boudreault C, Wilson AD, and Scheffler BE. 2012. Siricidae (Hymenoptera: Symphyta: Siricoidea) of the Western Hemisphere. Canadian Journal of Arthropod Identification 21: 1-305.
    )

May be confused with

Sirex obesus females can be distinguished from S. areolatus and the dark form of S. californicus by the length of the second hind tarsomeretarsomere:
a segment of the tarsus
and by the dense genagena:
the area of the head between the compound eye and clypeus; also called the cheek
pits. The males can be recognized by the black antennaeantenna:
the sensory organ emerging from the front of the head, usually between the compound eyes and above the clypeus; includes the flagellum, scape and pedicel
and the yellow-tinted fore wingfore wing:
the anterior wing of each pair of wings; usually the largest wing of the pair
(Schiff et al. 2012Schiff et al. 2012:
Schiff NM, Goulet H, Smith DR, Boudreault C, Wilson AD, and Scheffler BE. 2012. Siricidae (Hymenoptera: Symphyta: Siricoidea) of the Western Hemisphere. Canadian Journal of Arthropod Identification 21: 1-305.
).

Morphological and geographical variation

none recorded

Host associations

One specimen of S. obesus was collected from Pinus ponderosa (ponderosa pine) (Schiff et al. 2012Schiff et al. 2012:
Schiff NM, Goulet H, Smith DR, Boudreault C, Wilson AD, and Scheffler BE. 2012. Siricidae (Hymenoptera: Symphyta: Siricoidea) of the Western Hemisphere. Canadian Journal of Arthropod Identification 21: 1-305.
).

Life history

Female Sirex harbor symbiotic basidiomycete fungus in abdominal glands called mycangia. During oviposition, the site is inoculated with the fungus, which begins to decompose the surrounding wood. LarvaeLarva:
the immature stage of holometabolous insects
feed on the fungus, and in the process bore galleries through the wood (Johnson 1930Johnson 1930:
Johnson CW. 1930. On the variation and abundance of Sirex nitidus Harris. Psyche 37 (3): 281-282. https://doi.org/10.1155/1930/62786
, Schiff et al. 2012Schiff et al. 2012:
Schiff NM, Goulet H, Smith DR, Boudreault C, Wilson AD, and Scheffler BE. 2012. Siricidae (Hymenoptera: Symphyta: Siricoidea) of the Western Hemisphere. Canadian Journal of Arthropod Identification 21: 1-305.
).

Larvae are creamy white and grub-like in appearance with a dark head capsule. As with adults, larvaelarva:
the immature stage of holometabolous insects
possess a short dorsaldorsal:
of or on the top surface of the body or structure
horn on the posterior end of the body. The larvaelarva:
the immature stage of holometabolous insects
bore galleries into wood, feeding until pupation and subsequent emergence. Throughout this process, the larvaelarva:
the immature stage of holometabolous insects
use their horn to pack the tunnel behind them with sawdust. Emergence holes are perfectly circular. The fungal symbiont is carried in specialized organs in female larvaelarva:
the immature stage of holometabolous insects
that develop into the mycangia after metamorphosis (Schiff et al. 2012Schiff et al. 2012:
Schiff NM, Goulet H, Smith DR, Boudreault C, Wilson AD, and Scheffler BE. 2012. Siricidae (Hymenoptera: Symphyta: Siricoidea) of the Western Hemisphere. Canadian Journal of Arthropod Identification 21: 1-305.
).

The flight period of S. obesus is from late July through late September (Schiff et al. 2012Schiff et al. 2012:
Schiff NM, Goulet H, Smith DR, Boudreault C, Wilson AD, and Scheffler BE. 2012. Siricidae (Hymenoptera: Symphyta: Siricoidea) of the Western Hemisphere. Canadian Journal of Arthropod Identification 21: 1-305.
).

Distribution

World: North America

North America: Sirex obesus is only known from Arizona and New Mexico in the southwestern United States (Schiff et al. 2012Schiff et al. 2012:
Schiff NM, Goulet H, Smith DR, Boudreault C, Wilson AD, and Scheffler BE. 2012. Siricidae (Hymenoptera: Symphyta: Siricoidea) of the Western Hemisphere. Canadian Journal of Arthropod Identification 21: 1-305.
).

No specific locality data was available for mapping the range of this species at the time of publication.

  Sirex obesus  female lateral habitus; photo by J. Orr, WSDA

Sirex obesus female lateral habitus; photo by J. Orr, WSDA

  Sirex obesus  female dorsal habitus; photo by J. Orr, WSDA

Sirex obesus female dorsal habitus; photo by J. Orr, WSDA

  Sirex obesus  male lateral habitus; photo by H. Goulet, CNC

Sirex obesus male lateral habitus; photo by H. Goulet, CNC

  Sirex obesus  fore wing; photo by H. Goulet, CNC

Sirex obesus fore wing; photo by H. Goulet, CNC

  Sirex obesus  male antenna; photo by H. Goulet, CNC

Sirex obesus male antenna; photo by H. Goulet, CNC

  Sirex obesus  head; photo by H. Goulet, CNC

Sirex obesus head; photo by H. Goulet, CNC

  Sirex obesus  ovipositor pits; photo by H. Goulet, CNC

Sirex obesus ovipositor pits; photo by H. Goulet, CNC