Tethida

Taxonomy

Family: Tenthredinidae
Family common name: common sawflies
Subfamily: Blennocampinae
Tribe: Tomostethini
Genus: Tethida Ross, 1937
Subgenera: none

Background

The Tenthredinidae are the most species-rich family and are found throughout the world, in all continents but Antarctica. They are known as the “common sawflies.” They can generally be recognized by a cylindrical body and long, segmented antennaeantenna:
the sensory organ emerging from the front of the head, usually between the compound eyes and above the clypeus; includes the flagellum, scape and pedicel
. Otherwise, they come in a variety of colors, sizes, and forms (Goulet 1992Goulet 1992:
Goulet H. 1992. The genera and subgenera of the sawflies of Canada and Alaska: Hymenoptera. Symphyta. The insects and arachnids of Canada. Part 20. Agriculture Canada Publication.
).

Sawflies in the subfamily Blennocampinae have a diverse set of life histories and habits. Many species are restricted to subtropical and tropical regions, but the genus is still fairly species-rich in North America. Blennocampinae includes many sawflies that feed on ornamental and forestry crops. This subfamily can be recognized by wing venationvenation:
the network of veins on a wing
and bidentatebidentate:
having two teeth; often used in descrbing mandibles or tarsal claws
mandibles (Smith 1969dSmith 1969d:
Smith DR. 1969d. Nearctic Sawflies. I. Blennocampinae: Adults and larvae (Hymenoptera: Tenthredinidae). Technical Bulletin, U.S. Department of Agriculture 1397: 1-176.
).

Tethida is monotypicmonotypic:
describes having only one representative; ex. a genus that includes only one species
. Tethida barda is about 7 mm in length and mostly black with reddish coloring on the laterallateral:
of or towards the side of the body
thorax and darkened wings (Smith 1969dSmith 1969d:
Smith DR. 1969d. Nearctic Sawflies. I. Blennocampinae: Adults and larvae (Hymenoptera: Tenthredinidae). Technical Bulletin, U.S. Department of Agriculture 1397: 1-176.
).

Diversity

There is a single described extantextant:
in existence; opposite of extinct
species worldwide, and it is North American (Taeger et al. 2018Taeger et al. 2018:
Taeger A, Liston AD, Prous M, Groll EK, Gehroldt T, and Blank SM. 2018. ECatSymmdash;Electronic World Catalog of Symphyta (Insecta, Hymenoptera). Program version 5.0 (19 Dec 2018), data version 40 (23 Sep 2018). Senckenberg Deutsches Entomologisches Institut (SDEI), Muuml;ncheberg. https://sdei.de/ecatsym/ Accessed: 28 Jan 2020.
).

Diagnostic characteristics

Subfamily characters

  • angle of intersection of fore wingfore wing:
    the anterior wing of each pair of wings; usually the largest wing of the pair
    veins Cu1 and 1m-cu between 120°–150° (Goulet 1992Goulet 1992:
    Goulet H. 1992. The genera and subgenera of the sawflies of Canada and Alaska: Hymenoptera. Symphyta. The insects and arachnids of Canada. Part 20. Agriculture Canada Publication.
    )
  • fore wing fore wing:
    the anterior wing of each pair of wings; usually the largest wing of the pair
    veins 2A and 3A incomplete (Smith 1969dSmith 1969d:
    Smith DR. 1969d. Nearctic Sawflies. I. Blennocampinae: Adults and larvae (Hymenoptera: Tenthredinidae). Technical Bulletin, U.S. Department of Agriculture 1397: 1-176.
    )

Genus characters

  • basal basal:
    towards the base; closest to the body
    fifth of the clypeusclypeus:
    sclerotized area on the front of the head located between the antennal insertions and labrum
    evenly haired (Goulet 1992Goulet 1992:
    Goulet H. 1992. The genera and subgenera of the sawflies of Canada and Alaska: Hymenoptera. Symphyta. The insects and arachnids of Canada. Part 20. Agriculture Canada Publication.
    )
  • epicnemial area clearly outlined (Goulet 1992Goulet 1992:
    Goulet H. 1992. The genera and subgenera of the sawflies of Canada and Alaska: Hymenoptera. Symphyta. The insects and arachnids of Canada. Part 20. Agriculture Canada Publication.
    )
  • fore wing fore wing:
    the anterior wing of each pair of wings; usually the largest wing of the pair
    veins M and 1m-cu narrowed at apexapex:
    the end or most distal area of any structure
    , not parallel (Smith 1969dSmith 1969d:
    Smith DR. 1969d. Nearctic Sawflies. I. Blennocampinae: Adults and larvae (Hymenoptera: Tenthredinidae). Technical Bulletin, U.S. Department of Agriculture 1397: 1-176.
    )
  • hind wing hind wing:
    the posterior wing of each pair of wings
    cellcell:
    1. a membranous area of the wing between veins, 2. a small cavity or closed space
    M present (Smith 1969dSmith 1969d:
    Smith DR. 1969d. Nearctic Sawflies. I. Blennocampinae: Adults and larvae (Hymenoptera: Tenthredinidae). Technical Bulletin, U.S. Department of Agriculture 1397: 1-176.
    )
  • mesonotum mesonotum:
    the second segment of the dorsum of the thorax
    sparsely haired on central surface (Goulet 1992Goulet 1992:
    Goulet H. 1992. The genera and subgenera of the sawflies of Canada and Alaska: Hymenoptera. Symphyta. The insects and arachnids of Canada. Part 20. Agriculture Canada Publication.
    )
  • pulvilli on the first and second tarsomeres well developed (Goulet 1992Goulet 1992:
    Goulet H. 1992. The genera and subgenera of the sawflies of Canada and Alaska: Hymenoptera. Symphyta. The insects and arachnids of Canada. Part 20. Agriculture Canada Publication.
    )
  • tarsal claw tarsal claw:
    sharpened appendage emerging from the apex of the tarsus
    appearing simple but with very small inner tooth (Smith 1969dSmith 1969d:
    Smith DR. 1969d. Nearctic Sawflies. I. Blennocampinae: Adults and larvae (Hymenoptera: Tenthredinidae). Technical Bulletin, U.S. Department of Agriculture 1397: 1-176.
    )
  • pits not present on mesoscutellummesoscutellum:
    the anterior section of the scutellum
    and mesepisternummesepisternum:
    the ventral portion of the mesopleuron, located between the forecoxae and mid coxae
    (Goulet 1992Goulet 1992:
    Goulet H. 1992. The genera and subgenera of the sawflies of Canada and Alaska: Hymenoptera. Symphyta. The insects and arachnids of Canada. Part 20. Agriculture Canada Publication.
    )

May be confused with

Tethida can be confused with similar species in the subfamily Blennocampinae. It can be distinguished from most other genera by the developed pulvillipulvillus:
soft pads used for surface adhesion, located in sawflies on the first 4 segments of the tarsus
on basalbasal:
towards the base; closest to the body
tarsomeres and by the epicnemialepicnemium:
the anterior area of the mesepisternum
area, and from similar genus Tomostethus by the sparsely haired mesonotummesonotum:
the second segment of the dorsum of the thorax
and the basalbasal:
towards the base; closest to the body
area of the clypeusclypeus:
sclerotized area on the front of the head located between the antennal insertions and labrum
(Goulet 1992Goulet 1992:
Goulet H. 1992. The genera and subgenera of the sawflies of Canada and Alaska: Hymenoptera. Symphyta. The insects and arachnids of Canada. Part 20. Agriculture Canada Publication.
).

Exotic pest species of concern

none

Host associations

Tethida barda feeds on species of Fraxinus (ash) (Goulet 1992Goulet 1992:
Goulet H. 1992. The genera and subgenera of the sawflies of Canada and Alaska: Hymenoptera. Symphyta. The insects and arachnids of Canada. Part 20. Agriculture Canada Publication.
), including F. americana (white ash) (Schlesinger 1990Schlesinger 1990:
Schlesinger RC. 1990. Fraxinus americana L. white ash. In: Burns RM and Honkala BH, eds. Silvics of North America Volume 2, Hardwoods. Pp. 333-338.
) and F. pennsylvanica (green ash) (Brakie 2013Brakie 2013:
Brakie M. 2013. Plant guide for green ash ( Fraxinus pennsylvanica ). USDA Natural Resources Conservation Service, East Texas Plant Materials Center. Nacogdoches, Texas. 75964.
).

Life history

Tethida barda is a pest of ash trees known as the black-headed ash sawfly (Brakie 2013Brakie 2013:
Brakie M. 2013. Plant guide for green ash ( Fraxinus pennsylvanica ). USDA Natural Resources Conservation Service, East Texas Plant Materials Center. Nacogdoches, Texas. 75964.
). Females oviposit into the edge of the leaves in rows, leaving a blistered patterning on the leaf. After hatching, larvaelarva:
the immature stage of holometabolous insects
feed gregariously on the underside of the leaf, consuming it entirely. LarvaeLarva:
the immature stage of holometabolous insects
are white to light green with black head capsules (Osborne 1884Osborne 1884:
Osborne H. 1884. The ash saw-fly ( Selandria barda Say). The Canadian Entomologist 16 (8): 148-152.
).

Distribution

World: This genus is known only from North America (Taeger et al. 2010Taeger et al. 2010:
Taeger A, Blank SM, and Liston AD. 2010. World Catalog of Symphyta (Hymenoptera). Zootaxa 2580: 1-1064.
).

North America: Tethida barda occurs in the eastern part of North America, from as far north as New Brunswick and Saskatchewan, south to Florida and Texas (Smith 1969dSmith 1969d:
Smith DR. 1969d. Nearctic Sawflies. I. Blennocampinae: Adults and larvae (Hymenoptera: Tenthredinidae). Technical Bulletin, U.S. Department of Agriculture 1397: 1-176.
).

Map data from: GBIF.org (29 October 2019) GBIF Occurrence Download Tethida

Details about data used for maps can be found here.

  Tethidanbsp;bardanbsp; female lateral habitus; photo by J. Orr, WSDA

Tethida barda female lateral habitus; photo by J. Orr, WSDA

  Tethidanbsp;bardanbsp; male lateral habitus; photo by J. Orr, WSDA

Tethida barda male lateral habitus; photo by J. Orr, WSDA

  Tethidanbsp;bardanbsp; male face; photo by Q. Baine, WSDA

Tethida barda male face; photo by Q. Baine, WSDA