Phymatodes

Subfamily

Cerambycinae

Diagnosis - adult

Body length: 3–16 mm. 
Eyes: eye interommatidial setaeseta:
a sclerotized hair-like projection of the cuticle
absent, eye deeply emarginateemarginate:
notched at the margin
> half width or divided—max one row of facets between, eye ommatidial density fine. 
AntennaeAntenna:
in larval and adult insects, paired segmented appendages, borne one on each side of the head, functioning as sense organs and bearing a large number of sensilla
: antennal length reaches between basebase:
the part of any appendage or structure that is nearest the body
and end of elytraelytron:
the leathery forewing of beetles, serving as a covering for the hind wings, commonly meeting opposite elytron in a straight line down the middle of the dorsum in repose
or reaching/surpassing end of body, antennal flagellar segments elongateelongate:
much longer than wide
, scapescape:
the first proximal segment of the antenna
smooth/punctate at apexapex:
end of any structure distad to the base
, antennal scapescape:
the first proximal segment of the antenna
≥ segment 3 or segment 3 > scapescape:
the first proximal segment of the antenna
. 
Pronotumpronotum:
the upper and dorsal part of the prothorax
: pronotumpronotum:
the upper and dorsal part of the prothorax
shape transversetransverse:
broader than long
or subquadratesubquadrate:
 not quite a square
, pronotumpronotum:
the upper and dorsal part of the prothorax
lateral armature absent. 
Prosternum: prosternal process not dilated at apexapex:
end of any structure distad to the base
, procoxal cavities open posteriorly. 
ElytraElytron:
the leathery forewing of beetles, serving as a covering for the hind wings, commonly meeting opposite elytron in a straight line down the middle of the dorsum in repose
: elytral length reaching or close to end of abdomen, elytral apicesapex:
end of any structure distad to the base
rounded or truncatetruncate:
cut off squarely at the tip
, rarely with tooth or spinespine:
a protuberance with an acute (sharp) distal end
, elytral color black, brown, metallic, reddish, or orange, elytral color pattern present or absent. 
Legs: visible tarsomerestarsomere:
subdivision or article of the tarsus, usually numbering from two to five
: 4, femora clavateclavate:
thickening gradually toward the tip
, protibial spurs: 2, tarsal clawstarsal claw:
usually paired claws of the pretarsus, at the distal end of the leg
simple.

Slender, subcylindrical, rarely flattened. subquadratesubquadrate:
 not quite a square
,1-segmented and shorter than body in both sexes, second segment less than twice as long as broad; palpi very unequal. Pronotumpronotum:
the upper and dorsal part of the prothorax
rarely transversetransverse:
broader than long
, sides rounded, intercoxal process of prosternum short, narrow, pointed behind; mesosternummesosternum:
sternum of the mesothorax
broad, acutely triangular. Femora clavateclavate:
thickening gradually toward the tip
; first segment of posterior tarsitarsus:
the leg segment distal to the apex of the tibia, bearing the pretarsus; consists of one to five tarsomeres (including pretarsus)
slender, more than twice as long as broad (Linsley 1964Linsley 1964:
Linsley EG. 1964. The Cerambycidae of North America. Part V. Taxonomy and Classification of the Subfamily Cerambycinae, Tribes Callichromini Through Ancylocerini. University of California Publications in Entomology, Vol. 22. 197 pp.).

Similar genera

Eumichthus, Callidium, Meriellum

Differential diagnosis

Phymatodes differs from most Callidiini in the prosternal process ending between the procoxaeprocoxa:
coxa of the front leg 
. It can be differentiated from Callidium by the presence of a stridulatum and no large punctures on the mesonotum. Eumichthus looks similar but differs from Phymatodes by the second antennal segment more than twice as long as wide/about half as long as third, the final palpal segment triangularly dilated, and the male with the proleg tarsomerestarsomere:
subdivision or article of the tarsus, usually numbering from two to five
1 and 2 swollen.

Distribution

Holarctic, Indomalaya; P. (Phymatoderus) lividus introduced to USA, Argentina, Uruguay

Recorded host plants

Abies, Pinus, Larix, Picea, Pseudotsuga, Tsuga; Juniperus, Cupressus, libocedrus, Sequoia, Thuja, Chamaecyparis, Hesperocyparis macrocarpa (P. nitidus)

Remarks

29 spp. (Phymatodes), 1 sp. (Melasmetus), 4 spp. (Paraphymatodes), 9 spp. (Phymatodellus), 9 spp. (Phymatoderus), 10 spp. (Poecilium). Conifers: P. (Phymatoderus) abietinus, P. (P.) glabratus, P. (Phymatodes) aeneus, P. (P.) testaceus, P. (P.) hirtellus, P. (P.) dimidiatus, P. (P.) fulgidus, P. (P.) hardyi, P. (P.) maculicollis, P. (P.) nigerrimus, P. (P.) nitidus,. P. testaceus is Holarctic.

Synonyms

Merium Kirby, 1837

Phymatodina Casey, 1912

Microcallidium Casey, 1912

Taxonomy

Phymatodes Mulsant (not Dejean, 1834), 1839