Subfamily: Faboideae.
Phylogenetic Number: 3.10.26.
Tribe: Phaseoleae.
Subtribe: Diocleinae.
Species Studied - Species in Genus: 5 studied; 5 in genus.
Fruit: A legume; unilocular; 4.2–13 cm long; 0.5–1 cm wide; 0.2–0.3 cm thick; 2–9 times longer than wide to more than 9 times longer than wide; with deciduous androecial sheath; with deciduous corolla; with persistent calyx; with calyx shorter than fruit; without orifice formed by curving of fruit or fruit segments; straight, or curved (slightly); not plicate; not twisted; symmetrical, or asymmetrical; linear; with both sutures nearly straight; not inflated; compressed; without beak, or with beak; straight; with solid beak the same color and texture as fruit; tapered at apex to short tapered at apex; oblique with longitudinal axis of fruit; tapered at base, or short tapered at base; aligned with longitudinal axis of fruit; with the apex and base uniform in texture; coriaceous; seed chambers externally visible, or invisible; margin not constricted; margin without sulcus; margin plain, or embellished; margin with thickened sutural areas; wing(s) absent; nonstipitate; with all layers dehiscing; splitting along suture(s). Dehiscence of valves along both sutures; apical and down; active; with valves twisting. Replum invisible. Epicarp dull; monochrome; brown; with surface texture uniform; pubescent and indurate; with hairs erect; with 1 type of pubescence; velutinous, or sericeous; with pubescence golden, or white; with pubescence uniformly distributed; with simple hairs; pliable; with hair bases plain; eglandular; without spines; smooth; not veined; not tuberculate; not exfoliating; with cracks; cracking oblique to fruit length; without embedded tissue, much thicker than epicarp, running from base to apex. Mesocarp present; thin; surface not veined; 2-layered; without balsamic vesicles; without fibers; without reniform canals; with spongy layer over solid layer; coriaceous to chartaceous. Endocarp present; visible; dull; opaque; mottled; brown, or tan; with mottling over seed chambers; with brown overlay; scurfy; without adhering pieces of testa; septate; with septa thicker than paper, firm; with septa eglandular; chartaceous; exfoliating in part; remaining fused to mesocarp and epicarp; without wings; entire. Seed(s) 6–14; length transverse to fruit length; neither overlapping nor touching; in 1 series. Funiculus 0.5–6 mm long; of 1 length only; flattened; triangular. Aril present; dry; when dry rim-aril; entire to crenate; covering less than 1/2 of seed; without tongue (or flap) on lips of 2-lipped rim-aril, or with tongues (or flap) on lips of 2-lipped rim-aril; with 2 tongues or flaps, 1 on each lip of 2-lipped rim-aril; tan.
Seed: 7–8.8 mm long; 5–5.8 mm wide; 3.5–5 mm thick; not overgrown; not angular; symmetrical; oblong to ovate; terete to compressed; with surface smooth; without visible radicle and cotyledon lobes; without hilar sinus; without umbo on seed faces; without medial ridge on each face. Cuticle not exfoliating; not inflated; not wrinkled. Testa present; without pieces of adhering epicarp; not adhering to endocarp; free from endocarp; dull; not modified by a bloom; colored; mottled; with frequent mottles; reddish brown; with brown overlay (dark); glabrous; smooth; chartaceous. Pleurogram absent. Pseudopleurogram absent. Fracture lines absent. Rim absent. Wing(s) absent. Raphe not visible. Hilum present; visible; with faboid split; with the lips of the faboid split lighter colored than the rest of the hilum and therefore conspicuous; larger than punctiform; 4–5 mm long; with straight outline; linear; subapical to radicle tip; raised; within rim, or within halo; halo darker than testa; rim color darker than testa. Lens discernible; 0.7–1.5 mm long; with margins straight; linear; not in groove of raphe; confluent with hilum; flush; dissimilar color from testa; lighter than testa; tan; within halo; halo color darker than testa. Endosperm present; trace; not pluglike and not resembling tip of radicle; restricted to region of embryo; adnate to testa. Cotyledons smooth; both outer faces convex; both the same thickness; both more or less of equal length; not folded; margin entire 180 degrees from base of radicle; similar at apex; partially concealing radicle; split over radicle; without lobes; with the interface division terminating at base of radicle; without margins recessed; tan; inner face flat; glabrous on inner face. Embryonic axis right angled; oblique to length of seed; without a joint evident between the radicle and the cotyledons. Radicle differentiated from cotyledon; linear; lobe tip straight; oblique to cotyledons; centered between cotyledons; less than 1/2 length of cotyledons. Plumule well developed; glabrous.