Subfamily: Faboideae.
Phylogenetic Number: 3.10.25.
Tribe: Phaseoleae.
Subtribe: Diocleinae.
Species Studied - Species in Genus: 2 studied; 5 in genus.
Fruit: A legume; unilocular; 6–18 cm long; 1.3–2 cm wide; 0.3–0.5 cm thick; 2–9 times longer than wide; with deciduous androecial sheath; with deciduous corolla; with deciduous calyx; without orifice formed by curving of fruit or fruit segments; straight to curved (slightly); not plicate; not twisted; asymmetrical, or symmetrical; linear, or falcate; with both sutures parallelly curved; not inflated; compressed; without beak, or with beak; straight, or declined; with solid beak the same color and texture as fruit; aligned with longitudinal axis of fruit, or oblique with longitudinal axis of fruit; tapered at base; aligned with longitudinal axis of fruit; with the apex and base uniform in texture; coriaceous; seed chambers externally visible, or invisible; seed chambers with the raised seed chambers not torulose; margin not constricted; margin without sulcus; margin plain; wing(s) absent; nonstipitate; with all layers dehiscing; splitting along suture(s). Dehiscence of valves along both sutures; apical and down; active; with valves twisting. Replum invisible. Epicarp dull; monochrome; brown; with surface texture uniform; pubescent and indurate; with hairs erect; with 1 type of pubescence; puberulent, or velutinous; with pubescence golden to white; with pubescence uniformly distributed; with simple hairs; pliable; with hair bases plain; eglandular; without spines; not smooth; with elevated features; veined, or not veined; irregularly veined; not tuberculate; scurfy; not exfoliating; with cracks, or without cracks; cracking oblique to fruit length; without embedded tissue, much thicker than epicarp, running from base to apex. Mesocarp present; thin; 2-layered, or 3-layered; without balsamic vesicles; without fibers; without reniform canals; with solid layer over solid layer, or spongy layer over solid layer; with vitriol layer over 2 distinct solid layers, or solid layer over 2 distinct solid layers; coriaceous. Endocarp present; visible; dull; opaque; beneath seeds monochrome and mottled (between); tan; with mottling above and below seed chambers; with brown overlay, or gray overlay; beneath seeds smooth, scurfy, and cracked (the last two between seeds); without adhering pieces of testa; subseptate; with septa thin (tissue paper-like), flexible; with septa eglandular; chartaceous; exfoliating in part; separating from mesocarp; without wings; entire. Seed(s) 4–10; length transverse to fruit length; neither overlapping nor touching; in 1 series. Funiculus 0.5–2 mm long; of 1 length only; flattened; straight. Aril present; dry; when dry tongue-aril; entire; covering less than 1/2 of seed; tan.
Seed: 9 mm long; 3 mm wide; 4 mm thick; not overgrown; not angular; asymmetrical; nearly circular; compressed; with surface smooth; without visible radicle and cotyledon lobes; without hilar sinus; without umbo on seed faces; without medial ridge on each face. Cuticle not exfoliating; not inflated; not wrinkled. Testa present; without pieces of adhering epicarp; not adhering to endocarp; free from endocarp; dull; not modified by a bloom; colored; mottled; with infrequent mottles; brown; with brown overlay (dark); glabrous; smooth; coriaceous. Pleurogram absent. Pseudopleurogram absent. Fracture lines absent. Rim absent. Raphe not visible. Hilum present; visible; with faboid split; with the lips of the faboid split lighter colored than the rest of the hilum and therefore conspicuous; larger than punctiform; 2.5 mm long; with curved outline; elliptic; subapical to radicle tip; recessed; within rim; rim color darker than testa. Lens discernible; 0.8 mm long; with margins straight; linear; not in groove of raphe; confluent with hilum; flush; same color as testa; brown; not within corona, halo, or rim. Endosperm present; trace; not pluglike and not resembling tip of radicle; restricted to region of embryo; adnate to testa. Cotyledons smooth; both outer faces convex; both the same thickness; both more or less of equal length; not folded; margin entire 180 degrees from base of radicle; similar at apex; not concealing radicle; entire over radicle; without lobes; with the interface division terminating at base of radicle; without margins recessed; tan; inner face flat; glabrous on inner face. Embryonic axis right angled; oblique to length of seed; without a joint evident between the radicle and the cotyledons. Radicle differentiated from cotyledon; linear; lobe tip straight; oblique to cotyledons; centered between cotyledons; less than 1/2 length of cotyledons. Plumule well developed; glabrous.