Ahrens D, Zorn C, Dhoj Y, Keller S, Nagel P. 2007. Illustrated key to phytophagous scarabs of Nepal (Coleoptera, Scarabaeidae). Opuscula Biogeographica Baseleensia 5: 1-44.
Ahrens D. 2007. Taxonomic changes and an updated catalogue for the Palaearctic Sericini (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa 1504: 1-51. (accessed 2015) http://www.researchgate.net/publication/233751791_Taxonomic_changes_and_an_updated_catalogue_for_the_Palaearctic_Sericini_%28Coleoptera_Scarabaeidae_Melolonthinae%29
Allou K, Morin J, Kouassi P, N'klo F, Rochat D. 2006. Oryctes monoceros trapping with synthetic pheromone and palm material in Ivory Coast. Journal of Chemical Ecology 32: 1743-1754
Alm S. 1996. The oriental beetle. Turfgrass Trends 5 (7): 9-13.
Aloalii I, Masamdu R, Theunis W, Thistleton B. 1993. Prospects for biological control of taro beetles, Papuana spp. (Coleoptera: Scarabaeidae), in the South Pacific [pp. 66-70]. In: Proceedings of the Sustainable Taro Culture for the Pacific Conference. University of Hawaii Publishing, Honolulu, Hawaii, USA. (accessed 2015) http://www.ctahr.hawaii.edu/oc/freepubs/pdf/RES-140-17.pdf
Andresen E. 2005. Effects of season and vegetation type on community organization of dung beetles in a tropical dry forest. Biotropica 37: 291-300.
Anonymous. 1910. Report of the Entomologist. Hawaii Board of Commissioners of Agriculture and Forestry. Division of Forestry. Honolulu, HI: Hawaiian Gazette Company.
Anonymous. 1915. Minutes of the March 4th, 1915 meeting. Proceedings of the Hawaiian Entomological Society 3: 113–114. (accessed 2015) http://books.google.com/books?id=irMUAAAAYAAJ&pg=PA114
Anonymous. 1923. Report of the entomologist. Hawaii Board of Commissioners of Agriculture and Forestry. Division of Forestry. Honolulu, HI: Hawaiian Gazette Company.
Anonymous. 1952A. Proceedings of the Hawaiian Entomological Society for the year 1952. Proceedings of the Hawaiian Entomological Society 15: 1-20. (accessed 2015) http://hdl.handle.net/10125/14802
Anonymous. 1952B. Notes and exhibitions. Proceedings of the Hawaiian Entomological Society for the year 1951 14: 363. (accessed 2015) http://scholarspace.manoa.hawaii.edu/bitstream/handle/10125/16249/PHES14_343-370_Notes_Exhib_1951.pdf?sequence=1
Anonymous. 1953. Proceedings of the Hawaiian Entomological Society for the year 1952. Proceedings of the Hawaiian Entomological Society 15: 373-390. (accessed 2015) http://hdl.handle.net/10125/14802
Anonymous. 1954. Proceedings of the Hawaiian Entomological Society for the year 1954. Proceedings of the Hawaiian Entomological Society 15: 373-390. (accessed 2015) http://hdl.handle.net/10125/14847
Anonymous. 1957. List of intercepted plant pests, 1956 (pests recorded from July 1, 1955, through June 30, 1956). United States Department of Agriculture, Plant Quarantine Division. Hyattsville, Maryland, USA
Anonymous. 1961. Notes and exhibitions. Proceedings of the Hawaiian Entomological Society for the Year 1961. 18: 14. (accessed 2015) http://hdl.handle.net/10125/10843
Anonymous. 1964. Notes and exhibitions. Proceedings of the Hawaiian Entomological Society 18: 344-347. (accessed 2015) http://hdl.handle.net/10125/10885
Anonymous. 1967. Notes and exhibitions. Proceedings of the Hawaiian Entomological Society 19: 321. (accessed 2015) http://hdl.handle.net/10125/10932
Anonymous. 1969. List of intercepted plant pests, 1968 (pests recorded from July 1, 1967, through June 30, 1968). United States Department of Agriculture, Plant Quarantine Division. Hyattsville, Maryland, USA
Anonymous. 1973. Hawaii insect report. Cooperative Economic Insect Report issued by Plant Protection and Quarantine Programs, US Department of Agriculture 23: 676.
Anonymous. 1974. List of intercepted plant pests, 1972 (pests recorded from July 1, 1971, through June 30, 1972). United States Department of Agriculture, Plant Quarantine Division. Hyattsville, Maryland, USA
Anonymous. 1974. Notes and exhibitions. Proceedings of the Hawaiian Entomological Society 21: 315-317. (accessed 2015) http://hdl.handle.net/10125/11017
Anonymous. 1977. Trox spp. in Hawaii. Proceedings of the Hawaiian Entomological Society 22: 397-398. (accessed 2015) http://scholarspace.manoa.hawaii.edu/bitstream/handle/10125/11074/22_3proc.pdf?sequence=1
Anonymous. 1984. Sub-regional training course on methods of controlling diseases, insects, and other pests of plants in the south Pacific. Government Experimental Farm, Vaini, Tonga. October 4-12, 1982
Anonymous. 1986. Progress in upland rice research: Proceedings of the 1985 Jakarta conference. International Rice Research Institute, Manila, Philippines.
Anonymous. 1989. Notes and exhibitions. Proceedings of the Hawaiian Entomological Society 29: 10. (accessed 2015) http://scholarspace.manoa.hawaii.edu/bitstream/handle/10125/11236/29_proc.pdf?sequence=1
Anonymous. 1989. Toward insect resistant maize for the Third World: Proceedings of the 1987 International Symposium on Methodologies for Developing Host Plant Resistance to Maize Insects. CIMMYT, Mexico City, Mexico.
Anonymous. 2012. 2012 Prioritized Offshore Pest List. USDA APHIS. (accessed 2015) http://www.aphis.usda.gov/plant_health/plant_pest_info/pest_detection/downloads/farmbill/PrioritizedOffshorePestList.pdf
Anonymous. 2014. Destructive beetles found on Oahu coconut trees. Hawaii Department of Agriculture. (accessed May 21 2015). http://hdoa.hawaii.gov/blog/main/crb/
Anonymous. 2014. Invasive species of Japan: Protaetia pryeri oschimana. National Institute for Environmental Studies. (accessed April 25 2015). http://www.nies.go.jp/biodiversity/invasive/DB/detail/60520e.html
Anonymous. 2015. Exomala orientalis (oriental beetle). Invasive Species Compendium. Wallingford, UK, CABI International. (accessed 2015). http://www.cabi.org/isc/datasheet/5510
Anonymous. 2015. Distributions for Onthophagus nuchicornis. GBIF.org. (accessed 5 May 2015) http://www.gbif.org/species/1091565/distributions
Arakaki N, Nagayama A, Shimizu Y, Yara K. 2015. Suppression of green chafer Anomala albopilosa (Coleoptera: Scarabaeidae) populations by mass trapping with light traps. Journal of Applied Entomology and Zoology Online Publication DOI: 10.1007/s13355-015-0333-z
Arakaki N, Shimoji Y, Wakamura S. 2009. Camphor: an attractant for the cupreous polished chafer, Protaetia pryeri pryeri (Janson) (Coleoptera: Scarabaeidae). Applied Entomology Zoology 44: 621-625. (accessed: 2015) https://www.jstage.jst.go.jp/article/aez/44/4/44_4_621/_pdf
Arrow G. 1917. The fauna of British India including Ceylon and Burma. Coleoptera Lamellicornia Part II. (Rutelinae, Desmonychinae, and Euchirinae). Taylor and Francis.
Asquith A and Messing RH. 1993. Contemporary Hawaiian insect fauna of a lowland agricultural area on Kaua'i: implications for local and island-wide fruit fly eradication programs. Pacific Science 47: 1-16. (accessed 2015) http://scholarspace.manoa.hawaii.edu/bitstream/handle/10125/1742/v47n1-1-16.pdf?sequence=1
Bailey P. 2007. Pests of field crops and pastures: identification and control. CSIRO Publishing, Clayton, Australia. (accessed 2015) https://books.google.com/books?id=izWZ8-YP4hoC&pg=PA480&lpg=PA480&dq=%22sisyphus+spinipes%22&source=bl&ots=EhSKN3Whb1&sig=bW_SedRYBvu3C51h02nx7fl3YRE&hl=en&sa=X&ei=1PreVLrrI8qNyASpiYLQDA&ved=0CEMQ6AEwCA#v=onepage&q=%22sisyphus%20spinipes%22&f=false
Baker C. 1966. Larval systematics of the Troginae in North America with notes on biologies and life histories (Coleoptera: Scarabaeidea). PhD dissertation, Oregon State University. (accessed 2015) http://ir.library.oregonstate.edu/xmlui/bitstream/handle/1957/47471/BakerCharlesW1967.pdf?sequence=1
Barden S. 2011. Red imported fire ant influences on white grub populations and soil foraging characteristics in managed turf grass. PhD dissertation, Auburn University, Alabama. (accessed 2015) https://etd.auburn.edu/bitstream/handle/10415/2569/Final%20Thesis%204-29-2011.pdf?sequence=2
Barkhouse J and Ridsdill-Smith T. 1986. Effect of soil moisture on brood ball production by Onthophagus binodis Thunberg and Euonitcellus intermedius (Reiche) (Coleoptera: Scarabaeinae). Australian Journal of Entomology 25: 75-78. DOI: 10.1111/j.1440-6055.1986.tb01076.x
Barrett J. 1966. The biology and ecology of Lepidiota vogeli, a brown pasture scarab (family Melolonthidae) of the highlands of New Guinea. Papua New Guinea Agricultural Journal 18: 25-38.
Bedford G. 1974. Descriptions of the larvae of some rhinoceros beetles (Coleoptera: Scarabaeidae: Dynastinae) associated with coconut palms in New Guinea. Bulletin of Entomological Research. 63: 445–47
Bedford G. 1975. Observations on the biology of Xylotrupes gideon (Coleoptera: Scrabaeidae: Dynastinae) in Melanesia. Journal of the Australian Entomological Society 14: 213-216. (accessed 2015) http://onlinelibrary.wiley.com/doi/10.1111/j.1440-6055.1975.tb02028.x/pdf
Bedford G. 1980. Biology, ecology, and control of palm rhinoceros beetles. Annual Review of Entomology 25: 309-339.
Bedford G. 2015. Oryctes rhinoceros (coconut rhinoceros beetle). Invasive Species Compendium. Wallingford, UK, CABI International. (accessed 2015). http://www.cabi.org/isc/datasheet/37974
Bertone M. 2004. Dung beetles (Coleoptera: Scarabaeidae and Geotrupidae) of North Carolina cattle pastures and their implications for pasture improvement. MS thesis, University of North Carolina State University. (accessed 2015) http://repository.lib.ncsu.edu/ir/bitstream/1840.16/1952/1/etd.pdf
Bezborodov V, Aistova E, Rogatnykh D. 2011. Anthophilous Plastinchatousye beetles (Coleoptera, Scareabaeidae) Russian Far East. Amur Zoological Journal 3: 20-34.
Bezborodov V. 2014. Lamellicorn beetles (Coleoptera: Scarabaeoidea) of the Kuril Islands (Sakhalin Region, Russia): taxonomical structure, fauna, ecology and zoogeography. Caucasian Entomology Bulletin 10 (1): 33-46. (accessed 2015) http://www.zin.ru/animalia/coleoptera/pdf/bezborodov_2014_cauc_ent_bul.pdf
Bhawane G, Mamlayya A, Wagh S, Chaugule A. 2012. Diversity of white grub beetles and their host range from northern Western Ghats, Kolhapur District (MS) India. The Bioscan 7: 589-596.
Bhuiyan K and Nishigaki J. 1996. Oviposition of the adult cupreous chafer, Anomala cuprea Hope Coleoptera: Scarabaeidae, at different water contents of the ovipositing medium under laboratory conditions. Applied Entomology and Zoology 32: 431-436.
Blume R. 1981. Glaphyrocanthon viridis viridis (Beauvois): description of larva and notes on biology (Coleoptera : Scarabaeidae). The Coleopterists Bulletin 35: 235-238. (accessed 2015) http://www.jstor.org/stable/4007950
Blume R. 1984. Euoniticellus intermedius (Coleoptera: Scarabaeidae): description of adults and immatures and biology of adults. Environmental Entomology 13: 1064-1068. (accessed 2015) http://ee.oxfordjournals.org/content/ee/13/4/1064.full.pdf
Bouchard P. 2014. The Book of Beetles. Chicago, IL: University of Chicago Press.
Bourquin O. 2002. Invertebrates recorded from the Northern Mariana Islands status 2002. CNMI Invertebrate Collection Crees-Northern Marianas College, Saipan.
Bousquet Y (editor). 1991. Checklist of Beetles of Canada and Alaska. Agriculture Canada, Ottawa, Ontario, Canada. (accessed 2015) http://esc-sec.ca/aafcmonographs/checklist_of_beetles_of_canada_and_alaska.pdf
Boving A. 1945. Description of the larva and pupa of the scarab beetle Ancylonycha mindanaona (Brenske). Journal of the Washington Academy of Science 35: 13-15. (accessed 2015) http://biostor.org/reference/134138
Brach V. 1977. Larvae of Onthophagus polyphemi polyphemi and Onthophagus tuberculifrons Coleoptera Scarabaeidae. Bulletin Southern California Academy of Sciences 76: 66-68.
Britton E. 1962. A new species of Lepidiota (Coleoptera: Melolonthinae) affecting sugar cane in Queensland. Australian Journal of Entomology 2: 4-5.
Brown P and Daigneault A. 2014. Cost-benefit analysis of managing the Papuana uninodis (Coleoptera: Scarabaeidae) taro beetle in Fiji. Journal of Economic Entomology 107:1866-1877. (accessed 2015) http://www.bioone.org/doi/pdf/10.1603/EC14212
Bryan E. 1949. Economic insects of Micronesia; report of the Insect Control Committee for Micronesia, 1947-1948. Washington: National Research Council, Washington DC, USA.
Buss E. 2003. Sugarcane grub. University of Florida, Featured Creatures. (accessed 2015). http://entnemdept.ufl.edu/creatures/orn/turf/sugarcane_grub.htm
Buss E. 2006. Flight activity and relative abundance of phytophagous scarabs (Coleoptera: Scarabaeoidea) from two locations in Florida. Florida Entomologist 89:32-40. (accessed 2015) http://www.fcla.edu/FlaEnt/fe89p32.pdf
Camino-Lavín M, Jiminez-Perez A, Castrejon-Gomez V, Castrejon-Ayala F, Figueroa-Brito R. 1996. Performance of a new trap for melolonthine scarabs, root pests. Southwestern Entomologist 21: 325–330. [text in Spanish] (accessed 2015) http://sswe.tamu.edu/PDF/SWE_V21_N3_P325-330.pdf
Cartwright O and Gordon R. 1971. Insects of Micronesia Coleoptera: Scarabaeidae. Insects of Micronesia 17: 256-296.
Chandra K and Gupta D. 2013. Taxonomic studies on dung beetles (Coleoptera: Scarabaeidae, Geotrupidae, Hybosoridae) of Chhattisgarh, India. Munis Entomology & Zoology 8: 331-360. (accessed 2015) http://www.munisentzool.org/yayin/vol8/issue1/331-360.pdf
Chandra K. 2000. Inventory of scarabaeid beetles (Coleoptera) from Madhya Pradesh, India. Zoos' Print Journal 15: 359-362. (accessed 2015) http://www.zoosprint.org/zooprintjournal/2000/november/359-362.pdf
Chappell M. 1984. Thermoregulation and energetics of the green fig beetle (Cotinus [sic] texana) during flight and foraging behavior. Physiological Zoology 57: 581–589. (accessed 2015) http://www.jstor.org/stable/30155984
Chen R, Klein M, Li Q, Li Y. 2014. Mass trapping Popillia quadriguttata using Popillia japonica (Coleoptera: Scarabaeidae) pheromone and floral lures in northeastern China. Environmental Entomology 43: 77-81. (accessed 2015) http://ee.oxfordjournals.org/content/ee/43/3/774.full.pdf
Cherry R and Coale F. 1994. Oviposition of the sugarcane grub, Ligyrus subtropicus (Coleoptera:Scarabaeidae) in different soils. Journal of Economic Entomology 11: 345-348.
Cherry R. 1985. Seasonal phenology of white grubs (Coleoptera: Scarabaeidae) in Florida sugarcane fields. Journal of Economic Entomology 78: 787-789.
Choo H, Kaya H, Huh J, Lee D, Kim Hyeong H, Lee S, Choo Y, 2002. Entomopathogenic nematodes (Steinernema spp. and Heterorhabditis bacteriophora) and a fungus Beauveria brongniartii for biological control of the white grubs, Ectinohoplia rufipes and Exomala orientalis, in Korean golf courses. BioControl 47: 177-192. DOI: 10.1023/A:1014559729607
Cosner C. 2013. Invasive species pathway risk analysis for California. PhD dissertation, University of California, Davis, California. (accessed 2015) http://www.iscc.ca.gov/docs/reports/CISAC-Pathway-Report-July-2013-web.pdf
Cutler G and Rogers R. 1990. New record of the Asiatic garden beetle, Maladera castanea (Arrow), in Atlantic Canada. Journal of the Entomological Society of Ontario 140: 40-45. (accessed 2015) http://www.entsocont.ca/uploads/3/0/2/6/30266933/140_40_45.pdf
Davis A, Holloway J, Huijbregts H, Krikken J, Kirk-Spriggs A, Sutton A. 2001. Dung beetles as indicators of change in the forests of northern Borneo. Journal of Applied Ecology 38:592–616. DOI: 10.1046/j.1365-2664.2001.00619.x
Davis C and Krauss N. 1964. Recent introductions for biological control in Hawaii—IX. Proceedings of the Hawaiian Entomological Society 18:391-398. (accessed 2015) http://hdl.handle.net/10125/10890
Davis C and Krauss N. 1967. Recent introductions for biological control in Hawaii-XII. Proceedings of the Hawaiian Entomological Society 19:375-380. (accessed 2015) http://hdl.handle.net/10125/10937
Davis C. 1960. Recent introductions for biological control in Hawaii—V. Proceedings of the Hawaiian Entomological Society 17: 244-248. (accessed 2015) http://hdl.handle.net/10125/10814
Davis R. 2014. Who's in your compost?. Utah Pest News, Utah Plant Pest Diagnostic Laboratory and USU Extension 8: 6. (accessed 2015) http://utahpests.usu.edu/files/uploads/UtahPests-Newsletter-summer14.pdf
Dechambre R and Lachaume G. 2001. Oryctes. Les Coleopteres du Monde 27: 1-72. Hillside Books, Canterbury, UK.
DeNitto G, Cannon P, Eglitis A, Glaeser J, Maffei H, Smith S. 2015. Risk and pathway assessment for the introduction of exotic insects and pathogens that could affect Hawai‘i’s native forests. US Forest Service. (accessed 2015) http://www.fs.fed.us/psw/publications/documents/psw_gtr250/psw_gtr250.pdf
Diagne A, Story R, Hammond A. 2006. Adult Phyllophaga ephilida host plant feeding preference. Florida Entomologist 89: 391-395. (accessed 2015) http://journals.fcla.edu/flaent/article/view/75562/73220
Eckman L. 2015. Host plant feeding preferences of the adult Asiatic garden beetle, Maladera castanea Arrow (Coleoptera: Scarabaeidae). MS thesis, University of Connecticut. (accessed 2015) http://digitalcommons.uconn.edu/cgi/viewcontent.cgi?article=1784&context=gs_theses
Edmonds W. 1994. Revision of Phanaeus MacLeay, a New World genus of scarabaeine dung beetles (Coleoptera: Scarabaeidae, Scarabaeinae). Natural History Museum of Los Angeles County Contributions in Science 443: 1-105. http://centerforsystematicentomology.org/supplemental/Edmonds-1994-small.pdf
Edwards P and Aschenborn H. 1987. Patterns of nesting and dung burial in Onitis dung beetles: implications for pasture productivity and fly control. Journal of Applied Ecology 24: 837-851. (accessed 2015) http://www.jstor.org/stable/2403984
Edwards P. 2007. Introduced dung beetles in Australia 1967-2007. Landcare Australia, Sydeny, New South Wales, Australia. (accessed 2015) http://www.landcareonline.com.au/?page_id=2790
Elias S, Berman D, Alfimov A. 2000. Late Pleistocene beetle faunas of Beringia: where east met west. Journal of Biogeography 27: 1349–1363. (accessed 2015) http://www.researchgate.net/profile/Scott_Elias/publication/227667439_Late_Pleistocene_beetle_faunas_of_Beringia_where_east_met_west/links/00b4951ed3791450c6000000.pdf
Firake D, Deshmukh G, Behere N, Azad-Thakur S, Ngachan S, Rowland J. 2013. First report of elephant beetles in the genus Xylotrupes Hope (Coleoptera: Scarabaeidae) attacking guava. The Coleopterists Bulletin 67: 608-610.
Forgie S. 2009. Reproductive activity of Onthophagus granulatus Boheman (Coleoptera: Scarabaeinae) in New Zealand: implications for its effectiveness in the control of pastoral dung. New Zealand Entomologist 32: 76-84. DOI: 10.1080/00779962.2009.9722179
Fujiyama S, Kasuga S, Takahashi F. 1979. Studies on the self-regulation of life cycle in Anomala cuprea Hope (Coleoptera : Scarabaeidae). III. The effects of larval density and delayed hatching on the life cycle. Applied Entomology and Zoology 14: 231-240.
Fullaway O. 1921. Horn fly control. The Hawaiian Forester and Agriculturist: Quarterly Magazine of Forestry, Entomology, Plant Inspection and Animal Industry 18: 219-221. (accessed 2015) https://books.google.com/books?id=81jlAAAAMAAJ
Gaimari S (editor). 2005. California plant pest & disease report. California Department of Food and Agriculture, Plant Pest Diagnostics Branch 22: 52 (July 2002-July 2005).
García-Moll A. 2013. Adoretus sinicus (Chinese rose beetle). CABI Invasive Species Compendium. (accessed 8 April 2015). http://www.cabi.org/isc/datasheet/3282
Génier F. 2000. A new North American Ateuchus Weber (Coleoptera: Scarabaeidae, Scarabaeinae). The Coleopterists Bulletin 54: 341-346. (accessed 2015) http://dx.doi.org/10.1649/0010-065X(2000)054%5B0341:ANNAAW%5D2.0.CO;2
Goldstein PZ and DeSalle R. 2010. Integrating DNA barcode data and taxonomic practice: determination, discovery, and description. Bioessays 33: 135–147. [DOI 10.1002/bies.201000036]
Goodrich M. 1966. A revision of the genus Cotinis (Coleoptera: Scarabaeidae). Annals of the Entomological Society of America 59:550–568.
Gordon R and Skelley P. 2007. A monograph of the Aphodiini inhabiting the United States and Canada (Coleoptera: Scarabaeidae: Aphodiinae). Memoirs of the American Entomological Institute 79: 1-580.
Grebennikov V, Ballerio A, Ocampo F, Scholtz C. 2004. Larvae of Ceratocanthidae and Hybosoridae (Coleoptera: Scarabaeoidea): study of morphology, phylogenetic analysis and evidence of paraphyly of Hybosoridae. Systematic Entomology 29: 524–543. (accessed 2015) http://digitalcommons.unl.edu/cgi/viewcontent.cgi?article=1127&context=entomologypapers
Gressitt J. 1953. The coconut rhinoceros beetle (Oryctes rhinoceros) with particular reference to the Palau Islands. Bernice P. Bischop Museum, Bulletin 212: 157. (accessed 2015) http://guaminsects.myspecies.info/sites/guaminsects.myspecies.info/files/Gressitt%20-%201953.pdf
Gujarathi G and Pejaver M. 2014. Life cycle studies of Protaetia aurichalcea (Cetoniinae: Cetonidae: Scarabaeidae) in household biocompost. Species 10: 86-97. (accessed: 2015) http://www.researchgate.net/publication/264419351_Life_cycle_studies_of_Protaetia_aurichalcea_(CetoniinaeCetonidaeScarabaeidae)_in_household_biocompost
Gyeltshen A and Hodges A. 2005. Popillia japonica Newman (Insecta: Coleoptera: Scarabaeidae). University of Florida, Featured Creatures. (accessed 2015). http://entnemdept.ufl.edu/creatures/orn/beetles/japanese_beetle.htm
Habeck D. 1963. Description of immature stages of the Chinese rose beetle, Adoretus sinicus Burmeister (Coleoptera: Scarabaeidae). Proceedings of the Hawaiian Entomological Society 18: 251-258. (accessed 2015) http://scholarspace.manoa.hawaii.edu/bitstream/handle/10125/10873/18_251-258.pdf
Halffter G, Huerta C, Lopez-Portillo J. 1994. Parental care and offspring survival in Copris incertus Say, a sub-social beetle. Animal Behaviour 52: 133-139.(accessed 2015) http://www.researchgate.net/publication/240444845_Parental_care_and_offspring_survival_in_Copris_incertus_Say_a_sub-social_beetle
Hardy A. 1991. A catalog of the Coleoptera of America North of Mexico: family Scarabaeidae, subfamilies Rutelinae and Dynastinae. USDA Agriculture Handbook 529-34b. (accessed 2015) http://entomology.si.edu/coleoptera/ColeopteraCatalog/Scarabaeidae_Rutelinae&Dynastinae_529-34b.pdf
Harpootlian P. 2001. Scarab beetles (Coleoptera: Scarabaeidae) of South Carolina. Clemson University Public Service Publishing, Clemson, SC.
Harris R, Onaga K, Blume R, Roth J, Summerlin J. 1982. Survey of beneficial insects in undisturbed cattle droppings on Oahu, Hawaii. Proceedings of the Hawaiian Entomological Society 24: 91-95.
Hawkeswood T. 1985. Host plant record for Figulus regularis Westwood (Coleoptera: Lucanidae). The Coleopterist Bulletin 39: 200. (accessed 2015) http://www.calodema.com/freefiles/43.pdf
Hawkeswood T. 2014. A new adult host record for Figulus regularis Westwood, 1834 (Coleoptera: Lucanidae). Calodema 345: 1-2. (accessed 2015) http://docz.io/doc/285533/figulus-regularis-paper
Hayes W. 1917. Studies on the life history of Ligyrus gibbosus (Coleoptera). Journal of Economic Entomology 10: 253-261. (accessed 2015) https://books.google.com/books?id=CRv0AAAAMAAJ&pg=PA253&lpg=PA253&dq=ligyrus+gibbosus&source=bl&ots=8TMjOwRS1y&sig=o50KareSz26MTXX2Mc0ODmCx7xA&hl=en&sa=X&ei=d7fwVIb-BtWyyASXwIKwBA&ved=0CCgQ6AEwAg%23v=onepage&q=ligyrus%20gibbosus&f=false
Held D and Ray C. 2009. Asiatic garden beetle Maladera castanea (Coleoptera: Scarabaeidae) grubs found in damaged turf in Alabama. Florida Entomologist 92: 670-672. (accessed 2015) http://www.jstor.org/stable/25594598
Hession R, Arita L, Furutani S, Fukada M. 1994. Field observations on the mating behavior of the Chinese rose beetle, Adoretus sinicus Burmeister (Coleoptera: Scarabaeidae), in Hawaii. Journal of Hawaiian and Pacific Agriculture 5: 37-42.
Hinckley A. 1973. Ecology of the coconut rhinoceros beetle, Oryctes rhinoceros (L.) (Coleoptera: Dynastidae). Biotropica 5: 111-116.
Hiramatsu A, Sakamaki Y, Kusigemati K. 2001. A list of pest-insects feeding on the kenaf in Kagoshima City with seasonal abundance of some major pest-insects. Bulletin of the Faculty of Agriculture, Kagoshima University 51: 1-7.
Hongo Y. 2003. Appraising behavior during male-male interaction in the Japanese horned beetle Trypoxylus dichotomus septentrionalis (Kono). Behavior 140: 501-517. (accessed 2015) http://www.jstor.org/stable/4536041
Hongo Y. 2006. Bark carving behavior of the Japanese horned beetle Trypoxylus dichotomus septentrionalis (Coleoptera: Scarabaeidae). Journal of Ethology 24: 201-204. (accessed 2015) http://link.springer.com/content/pdf/10.1007%2Fs10164-006-0202-x.pdf
Houston W, Feehan J, Runko S. 1982. Methods for harvesting large numbers of dung beetles (Coleoptera: Scarabaeidae). Journal of the Australian Entomological Society 21: 217–219. DOI: 10.1111/j.1440-6055.1982.tb01798.x
Howarth F. 1985. Impacts of alien land arthropods and mollusks on native plants and animals in Hawaii [pp. 149-179]. In: Hawaii's Terrestrial Ecosystems: Preservation and Management. University of Hawaii Publishing, Honolulu, Hawaii, USA. (accessed 2015) http://www.hear.org/books/hte1985/pdfs/hte1985howarth.pdf
Howden H and Cartwright O. 1963. Scarab beetles of the genus Onthophagus Latreille north of Mexico. Proceedings of the United States National Museum 114: 1-135.
Howden H. 2008. In past years - XIII 1970–1974. Scarabs Newsletter 30: 15–21.
Huerta C, García-Hernández M. 2013. Nesting behavior of Onthophagus incensus Say, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae). The Coleopterists Bulletin 67: 161-166. DOI: 10.1649/0010-065X-67.2.161
Huerta C, Martinez I, García-Hernández M. 2010. Preimaginal development of Onthophagus incensus Say, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae). The Coleopterists Bulletin 64: 365-371. DOI: 10.1649/0010-065X-64.4.365
Hull R, Alaouna M, Khanyile L, Bryne M, Ntwasa M. 2013. Lifestyle and host defense mechanisms of the dung beetle, Euoniticellus intermedius: the toll signaling pathway. Journal of Insect Science 13: 1-25. (accessed 2015) http://www.ncbi.nlm.nih.gov/pmc/articles/PMC4011371/pdf/031.013.10801.pdf
Ijima K, Takeuchi M. 2007. Life history of Protaetia orientalis (Coleoptera: Scarabaeidae) in the Kanto District; and food resource plants of P. orientalis in Japan. Japanese Agricultural Science, Tokyo University of Agriculture 52: 89–96.
Jackson TA, Klein MG. 2006. Scarabs as pests: a continuing problem. The Coleopterists Bulletin 60 (5): 102-119.
Jameson M, Oishi D, Ratcliffe B, McQuate G. 2009. Two additional invasive scarabaeoid beetles (Coleoptera: Scarabaeidae: Dynastinae) in Hawaii. Proceedings of the Hawaiian Entomological Society 41: 25–30. (accessed 2015) http://digitalcommons.unl.edu/cgi/viewcontent.cgi?article=1146&context=entomologypapers
Jameson M, Ratcliffe B. 2002. Introduction. Series Scarabaeiformia Crowson 1960 (=Lamellicornia), superfamily Scarabaeoidea Latreille 1802, pp. 1-5. In, Arnett, Thomas, Skelley, and Frank (eds.), American Beetles. Polyphaga: Scarabaeoidea through Curculionoidea. Volume 2. CRC Press, New York.
Joy P, Anjana R, Soumya K. 2013. Pests of pineapple and their management. Pineapple Research Station, Kerala Agricultural University, India. (accessed 2015) http://prsvkm.kau.in/sites/default/files/documents/pests_of_pineapple_and_thier_management.pdf
Karimbumkara S (editor). 2013. Oniticellus cinctus. Encyclopedia of Life. (accessed 14 April 2015) http://eol.org/pages/143000/overview
Katino K, Seki N, Chiba M. 2004. Larval nutritional environment determines adult size in Japanese horned beetles Allomyrina dichotoma. Ecological Research 19: 663-668. DOI: 10.1111/j.1440-1703.2004.00681.x
Kato Y, Hara M, Sawada M. 1980. Occurrence of cupreous chafers, Anomala cuprea Hope, in the peanut field in Chiba Prefecture and their chemical control. Proceedings of the Kanto-Tosan Plant Protection Society 27: 130-131. (in Japanese) (accessed 2015) http://dx.doi.org/10.11337/ktpps1954.1980.130
Kim C, Lee S, Go M, Park K. 2002. Ecological characteristics of Protaetia orientalis submarmorea (Burmeister) (Coleoptera: Cetoniidae). Korean Journal of Applied Entomology 41: 43-47. [in Korean]
Kim J, Lee O. 1997. Taxonomy of Korean Sericinae (Coleoptera: Melolonthidae) II, genus Maladera. Insecta Koreana 14: 119-136. (accessed 2015) http://www.kangwon.ac.kr/~cisweb/ik14-6.html
Kim M, Lee H, Park H. 2003. A taxonomic study of immature stage in three species of the genus Protaetia Burmeister (Coleoptera: Scarabaeoidea, Cetoniidae) from Korea. Entomological Research, 33: 231-236. DOI: 10.1111/j.1748-5967.2003.tb00074.x
Kim S, Kim J. 2010. Review of family Lucanidae (Insecta: Coleoptera) in Korea with the description of one new species. Entomological Research 40: 55-81. DOI: 10.1111/j.1748-5967.2009.00263.x
Klein M. 2008. Popillia japonica (Japanese beetle). Invasive Species Compendium. Wallingford, UK, CAB International. Available from http://www.cabi.org/isc/datasheet/43599 (accessed April 20 2015). http://www.cabi.org/isc/datasheet/43599
Klemperer H. 1983. Subsocial behaviour in Oniticellus cinctus Coleoptera, Scarabaeidae: effect of the brood on parental care and oviposition. Physiological Entomology 84: 393-402.
Kobayashi H and Matsumoto T. 2011. Atlas of Japanese Scarabaeoidea. Volume 2 Phytophagous Group I. Roppon-Ashi Entomological Books, Tokyo, Japan.
Kohlmann B, Cano E, Delgado L. 2003. New species and records of Copris (Coleoptera: Scarabaeidae; Scarabaeinae) from Central America. Zootaxa 167: 1-16. (accessed 2015) http://www.mapress.com/zootaxa/2003f/zt00167.pdf
Kostromytska O and Buss E. 2011. Tomarus subtropicus (Coleoptera: Scarabaeidae) larval feeding habits. Florida Entomologist 94: 164-171.
Krajcik M. 1998. Cetoniidae of the world. Catalogue-Part I (Coleoptera: Cetoniidae). Typos Studio Most, Most, Czech Republic.
Krajcik M. 1999. Cetoniidae of the world. Catalogue-Part 2 (Coleoptera: Cetoniidae). Typos Studio Most, Most, Czech Republic.
Krajcik M. 2006. Checklist of the Scarabaeoidea of the world. 1. Scarabaeinae (Coleoptera: Scarabaeidae: Scarabaeinae). Animma.x. Supplement 3. Milan Kracjik, Plzen, Czech Republic.
Krajcik M. 2008. Checklist of the Scarabaeoidea of the world. 2. Rutelinae (Coleoptera: Scarabaeidae: Rutelinae), second edition. Animma.x. Supplement 4. Milan Kracjik, Plzen, Czech Republic.
Krell F, Hangay G. 1998. The African rhinoceros beetle Temnorhynchus retusus (Fabricius) established in eastern Australia (Coleoptera: Scarabaeidae: Dynastinae). Australian Journal of Entomology 37: 312–314. (accessed 2015) https://www.dmns.org/media/362198/56-austjent1998.pdf
Krell F. 1993. Phylogenetisch-systematische Revision des Genus Temnorhynchus Hope, 1837 (Coleoptera: Scarabaeoidea: Melolonthidae: Dynastinae: Pentodontini). 1. Teil: Phylogenetische Analyse, mit Anmerkunge zur phylogenetisch-sytematischen Methodologie. Beiträge zur Entomologie 43: 237–318.
Krikken J. 1977. Notes on African Onitini, mainly from southeastern Kenya (Coleoptera: Scarabaeidae). Zoologische Mededelingen: 141-170. (accessed 2015) http://www.repository.naturalis.nl/record/318134
Krischik V, Davidson J. 2004. Oriental beetle. Integrated Pest Management of Midwest Landscapes. University of Minnesota. Minnesota Agriculture Experiment Station, St. Paul, MN.
Krischik V. 2011. Japanese beetle management in Minnesota. Minnesota Department of Agriculture, University of Minnesota Extension Service. (accessed 2015). http://www.extension.umn.edu/garden/insects/find/japanese-beetles/
Krivolutskaya G. 1973. Entomofauna of the Kuril Islands: Principal Features and Origin. Izdatel'stvo Nauka, Leningrad Division. 315 pp. Translated from Russian by E Urdang. (accessed 2015). http://www.burkemuseum.org/static/okhotskia/ikip/Results/publications/entobook/title.htm
Krombein, K. 1949. The aculeate Hymenoptera of Micronesia I. Scoliidae, Mutillidae, Pompilidae, and Sphecidae. Proceedings of the Hawaiian Entomological Society 13: 367-410. (accessed 2015) https://scholarspace.manoa.hawaii.edu/handle/10125/16191
Krushelnycky P, Loope L, Gillespie R. 2007. Inventory of arthropods of the west slope shrubland and alpine ecosystems of Haleakala National Park. University of Hawaii at Manoa, Department of Botany. Technical Report. DOI: http://hdl.handle.net/10125/29487
Kuniata L and Young G. 1992. The biology of Lepidiota reuleauxi Brenske (Coleoptera: Scarabaeidae), a pest of sugarcane in Papua New Guinea. Journal of the Australian Entomological Society 31: 339-343.
Lal S, Moxon J, Autar M, Milner R, Hunter D, Hazelman S, Atumurirava F, Kurika K, Vasuidreketi T. 2008. Taro beetle management in Papua New Guinea and Fiji. Final Project Report, ACIAR Project Number CS2/2000/044. (accessed 2015) http://www.spc.int/lrd/publications/doc_download/1133-taro-beetle-report.pdf
Lansberge J. 1886. Scarabaeides, Buprestides, et Cerembycides de L'Afrique occidentale. Notes from the Leyden Museum. 8: 79-108.
Latchininsky A. 2008. Grasshopper outbreak challenges conservation status of a small Hawaiian island. Journal of Insect Conservation 12: 343–357. (accessed 2015) http://www.uwyo.edu/esm/faculty-and-staff/latchininsky/documents/2008-latchininsky-jico.pdf
Leblanc L, Graham S, Pohlman K, Fujita B. 2013. Protaetia orientalis (Coleoptera: Scarabaeidae) attracted to methyl eugenol fruit fly lure. Proceedings of the Hawaiian Entomological Society 45: 45–48. (accessed 2015) http://scholarspace.manoa.hawaii.edu/bitstream/handle/10125/31002/PHES45_45-48.pdf?sequence=1
Lee D, Choo H, Chung J, Lee S, Sagong Y. 2002. Hosts plants of Popillia quadriguttata (Coleoptera: Scarabaeidae). Korean Journal of Applied Entomology 41: 15-19.
Lee D, Choo H, Smitley D, Lee S, Shin H, Kaya H, Park C, Park J. 2007. Distribution and adult activity of Popillia quadriguttata (Coleoptera: Scarabaeidae) on golf courses in Korea. Journal of Economic Entomology 100: 103-109. DOI: doi.org/10.1603/0022-0493(2007)100[103:DAAAOP]2.0.CO;2
Liebherr J, Casale A, Rewin T, Nall G. 2010. On the provenance of Boheman's "Eugenies Resa" Carabidae (Coleoptera) allegedly described from Hawaii. The Coleopterists Society 64: 221-229. DOI: http://dx.doi.org/10.1649/0010-065X-64.3.221.7
Litsinger J, Barrion A, Soekarna D. 1987. Upland rice insect pests: their ecology, importance, and control. International Rice Research Institute. Manila, Philippines.
Litsinger J, Libetario E, Barrion A. 2002. Population dynamics of white grubs in the upland rice and maize environment of Northern Mindanao, Philippines. International Journal of Pest Management 48: 239-260. DOI: 10.1080/09670870210131861.
Look W. 1974. Efficacy of Hawaii's plant quarantine program. Proceedings of the Hawaiian Entomological Society 21: 413-418. (accessed 2015) http://hdl.handle.net/10125/11028
Losey JE, Vaughan M. 2006. The economic value of ecological services provided by insects. Bioscience 56 (4): 311-323.
Lumaret J. 1979. Contribution a l'etude des larves des Scarabaeidae. 7. Les larves de la tribu des Oniticellini (Coleoptera). Annales de la Societe Entomologique de France 15: 553-574.
MacQueen A and Beirne B. 1975. Dung burial activity and fly control potential of Onthophagus nuchicornis (Coleoptera: Scarabaeinae) in British Columbia. Canadian Journal of Plant Science 55: 961-967. DOI: 10.4141/cjps75-152
MacRae T and Penn S. 2001. Additional records of adventive Onthophagus Latreille (Coleoptera: Scarabaeidae) in North America. The Coleopterists Bulletin, 55: 49-50. (accessed, 2015) http://www.bioone.org/doi/full/10.1649/0010-065X%282001%29055%5B0049%3AAROAOL%5D2.0.CO%3B2
Maehler K. 1953. Beetles intercepted in quarantine. Proceedings of the Hawaiian Entomological Society 15: 13.
Mansfield-Aders W. 1920. Insects injurious to economic crops in the Zanzibar Protectorate. Bulletin of Entomological Research 10: 145-155. DOI: http://dx.doi.org/10.1017/S0007485300043947
Markin G and Yoshioka E. 1998. Biological control of the horn fly, Haematobia irritans L., in Hawai'i (Diptera: Muscidae). Proceedings of the Hawaiian Entomological Society 33: 43-50. (accessed 2015) http://scholarspace.manoa.hawaii.edu/bitstream/handle/10125/16301/PHES33_43-50.pdf?sequence=1
Marler T and Moore A. 2011. Military threats to terrestrial resources not restricted to wartime: a case study from Guam. Journal of Environmental Science and Engineering 5: 1198-1214. (accessed 2015) http://www.wptrc.org/userfiles/file/Marler/military%20buildup%20reprint.pdf
Marler T and Muniappan R. 2006. Pests of Cycas micronesica leaf, stem, and male reproductive tissues with notes on current threat status. Micronesica 19: 1-9. (accessed 2015) http://www.cerambycoidea.com/titles/marlermuniappan2006.pdf
Matthews E. 1972. A revision of the Scarabaeine dung beetles of Australia. I. Tribe Onthophagini. Australian Journal of Zoology Supplementary Series 19: 3-330.
Mau R and Kessing J. 1991. Adoretus sinicus. University of Hawaii Publishing, Honolulu, Hawaii, USA. (accessed 2015). http://www.extento.hawaii.edu/kbase/crop/Type/adoretus.htm
McCormack G. 2007. Species page: mango-flower beetle. Cook Islands Biodiversity Database, Cook Islands Natural Heritage Trust. (accessed 2015). http://cookislands.bishopmuseum.org/species.asp?id=14112
McQuate G and Jameson M. 2011. Sex determination in the Chinese rose beetle, Adoretus sinicus, and overview of Adoretus species of biosecurity concern. Journal of Insect Science 64: 1-18. (accessed 2015) http://dx.doi.org/10.1673/031.011.6401
Mitsuhashi W, Asano S, Miyamoto K, Wada S. 2014. Further research on the biological function of inclusion bodies of Anomala cuprea entomopoxvirus, with special reference to the effect on the insecticidal activity of a Bacillus thuringiensis formulation. Pest Management Science 70: 46-54. DOI: 10.1002/ps.3521
Molet T. 2013. CPHST pest datasheet for Oryctes rhinoceros. USDA-APHIS-PPQ-CPHST. (accessed April 9 2015). https://caps.ceris.purdue.edu/webfm_send/2206
Monteith G. 2011. Rhinoceros beetles. Queensland Museum Fact Sheet. (accessed 2015) http://www.qm.qld.gov.au/Learning+Resources/~/media/Documents/Learning%20resources/QM/Resources/Fact%20Sheets/fact-sheet-rhinoceros-beetles.pdf
Moore A. 2007. Assessment of the Rhinoceros Beetle Infestation on Guam. (accessed May 19 2015) http://guaminsects.myspecies.info/assessment-rhinoce-beetle-infestation-guam
Moore A. 2009. Guam coconut rhinoceros beetle (CRB) eradication program semi-annual progress report. University of Guam Cooperative Extension Service. 13 pp.
Moore A. 2010. Invasive insects on Guam. Micronesia Biosecurity Plan Meeting, Guam 2010. (accessed 2015). http://aliens.guaminsects.net/sites/default/files/GuamInvasiveInsectsMBP20100120.pdf
Moore A. 2012. Guam as a source of new insects for Hawaii. Pacific Entomology Conference, Portland, Oregon 2012. (accessed 2015). http://guaminsects.myspecies.info/sites/guaminsects.myspecies.info/files/GuamHawaiiBugConnection.pdf
Mora-Aguilar E and Delgado L. Two new species of Copris (Coleoptera: Scarabaeidae: Scarabaeinae) from Mexico, with a key based on major males to the remotus species complex. Annals of the Entomological Society of America 108: 875-880. DOI: http://dx.doi.org/10.1093/aesa/sav063 (accessed 2015)
Mori H and Chiba S. 2009. Sociality improves larval growth in the stag beetle Figulus binodulus (Coleoptera: Lucanidae). European Journal of Entomology 106: 379-383. (accessed 2015) http://www.eje.cz/pdfs/eje/2009/03/09.pdf
Muniappan R. 2012. Arthropod Pests of Horticultural Crops in Tropical Asia. CABI Publishing, Boston, USA.
Myers D, Schillinger J, Patton T, Sorensen K, Tubene S. 2003. On-farm sweet potato summer oriental beetle surveys for the development of an IPM action threshold for the prevention of juvenile white grub damage to mature roots. University of Maryland Extension, Glen Burnie, MD. (accessed 2015). https://extension.umd.edu/sites/default/files/_docs/VEGWKR%20Sweet%20Potato%20Study.pdf
Nafus D and Schreiner L. 1989. Biological control activities in the Mariana Islands from 1911 to 1988. Micronesica 22: 65-106. (accessed 2015) http://guaminsects.myspecies.info/sites/guaminsects.myspecies.info/files/Nafus1989a.pdf
Nair S, Prameela P, Suma A, Cherian K, Babu K. 2001. A new record of Xylotrupes gideon (L.) on okra (Abelmoschus esculentus [L.]). Insect Environment: 7: 71.
Nakao H and Funasaki G. 1976. Introductions for biological control in Hawaii, 1974. Proceedings of the Hawaiian Entomological Society 22: 329-331. (accessed 2015) http://hdl.handle.net/10125/11067
Nakao H and Funasaki G. 1979. Introductions for biological control in Hawaii: 1975 & 1976. Proceedings of the Hawaiian Entomological Society 23: 125-128. (accessed 2015) http://hdl.handle.net/10125/11098
Nakao H, Funasaki GY, Davis CJ. 1975. Introductions for biological control in Hawaii, 1973. Proceedings of the Hawaiian Entomological Society 22: 109-112.
Navarrete-Heredia J. 2012. New state records for Coleoptera in northern Mexico. The Coleopterists Bulletin 66: 74-65. http://dx.doi.org/10.1649/072.066.0118
Nishida G (editor). 2002. Hawaiian terrestrial arthropod checklist, fourth edition. Bishop Museum Technical Report 22: 1-313.
Nishida G and Beardsley J. 2002. A review of the insects and related arthropods of Midway Atoll. Bishop Museum Occasional Papers 68: 25–69. (accessed 2015) http://hbs.bishopmuseum.org/pdf/op68-04.pdf
Nitto M and Taohibana K. 1955. Decision of the number of instar and generation by means of measurements on the larval head, of May-beetle. Journal of the Japanese Forestry Society 37: 326-333. [In Japanese]
Noriega J, Horgan F, Larsen T, Valencia G. 2010. Records of an invasive dung beetle species, Digitonthophagus gazella (Fabricius, 1787) (Coleoptera: Scarabaeidae), in Peru. Acta Zoológica Mexicana 26: 451-456. (accessed 2015) http://www.scielo.org.mx/pdf/azm/v26n2/v26n2a9.pdf
Novelo E, Delfín-González H, Moron M. 2007. Copro-necrophagous beetle (Coleoptera: Scarabaeidae) diversity in an agroecosystem in Yucatan, Mexico. Revista de Biología Tropical 55: 83-99. (accessed 2015) http://www.scielo.sa.cr/scielo.php?pid=S0034-77442007000100009&script=sci_arttext&tlng=pt
Oakley R. 1945. Preliminary life-history studies in Guam of the scarab beetle Ancylonycha mindanaona (Brenske). Journal of the Washington Academy of Science 35: 7-12. (accessed 2015) http://biostor.org/reference/134137
Ocampo F. 2002. Hybosorids of the United States and expanding distribution of the introduced species Hybosorus illigeri (Coleoptera: Scarabaeoidea: Hybosoridae). Annals of the Entmological Society of America 95: 316-322. (accessed 2015) http://digitalcommons.unl.edu/cgi/viewcontent.cgi?article=1122&context=entomologypapers
Ohaus F. 1935. Checklist of the Rutelinae (Coleoptera, Scarabaeidae) of Oceania. Bernice P. Bishop Museum Occasional Papers 11: 3-9.
Ong X, Chua S, Potts M. 2013. Recent records of the dung beetle Carthsius molossus (Coleoptera: Scarabaeiidae) in Singapore. Nature in Singapore 6: 1-6. (accessed 2015) http://lkcnhm.nus.edu.sg/nis/bulletin2013/2013nis001-006.pdf
Ortega-Martinez I, Moreno C, Sanchez-Rojas G, Barragan F. 2014. The role of Canthon humectus hidalgoensis (Bates) (Coleoptera: Scarabaeidae) in dung removal from a cattle pasture. The Coleopterists Bulletin 68 (4): 656-658. (accessed 2015) http://dx.doi.org/10.1649/0010-065X-68.4.656
Osborn, HT. 1920. A note on Apterocyclus (Coleoptera, Lucanidae). Proceedings of the Hawaiian Entomological Society 4: 375-376. (accessed 2015) http://hdl.handle.net/10125/15668
Palestrini C, Pavan G, Zunino M. 1990. Acoustic signals in Copris incertus Say (Coleoptera Scarabaeidae Coprinae). Ethology Ecology & Evolution 3: 143-146. DOI: 10.1080/03949370.1991.10721928
Park S, Lim H, Kim D. 2014. A survey on insect diversity of Baengnyeongdo, Korea. Journal of Asia-Pacific Biodiversity 7: 268-280. (accessed 2015) http://dx.doi.org/10.1016/j.japb.2014.04.006
Paulsen M and Hawks D. 2014. A review of the primary types of the Hawaiian stag beetle genus Apterocyclus Waterhouse (Coleoptera, Lucanidae, Lucaninae), with the description of a new species. ZooKeys 433: 77–88. DOI: 10.3897/zookeys.433.8022
Pemberton C. 1954. Report for 1949-1954, invertebrates consultant committee for the Pacific. The Pacific Science Board, National Academy of Sciences, National Research Council, Washington, D.C., USA.
Pemberton C. 1963. Insect pests affecting sugar cane plantations within the Pacific, Proceedings of the 11th Congress of International Society of Sugar Cane Technologists, Mauritius. (accessed 2015) http://www.issct.org/pdf/proceedings/1962/1962%20Pemberton%20Insect%20Pests%20Affecting%20Sugar%20Cane%20Plantations.pdf
Pemberton C. 1964. Highlights in the history of entomology in Hawaii 1778-1963. Pacific Insects 6: 689-729. (accessed 2015) http://hbs.bishopmuseum.org/fiji/pdf/pemberton1964.pdf
Pena J, Sharp J, Wysoki M. 2002. Tropical Fruit Pests and Pollinators: Biology, Economic Importance, Natural Enemies, and Control. CABI Publishing, Boston, USA.
Perris E. 1887. Larves de Coléoptères. Annales de la Société Linnéenne de Lyon 23: 109. [in French]
Peterson G. 1956. An annotated checklist of parasites and predators introduced into Guam during the years 1950-1955. Proceedings of the Hawaiian Entomological Society 16: 199-202. (accessed 2015) http://hdl.handle.net/10125/14893
Plaistow S, Tsuchida K, Tsubaki Y, Setsuda K. 2005. The effect of a seasonal time constraint on development time, body size, condition, and morph determination in the horned beetle Allomyrina dichotoma L. (Coleoptera: Scarabaeidae). Ecological Entomology 30: 692-699. DOI:10.1111/j.0307-6946.2005.00740.x/full
Potter D and Held D. 2002. Biology and management of Japanese beetles. Annual Review of Entomology 47: 175-205.
Price D and May M. 2009. Behavioral ecology of Phanaeus dung beetles (Coleoptera: Scarabaeidae): review and new observations. Acta Zoológica Mexicana 25: 211-238. (accessed 2015) http://www.scielo.org.mx/scielo.php?pid=S0065-17372009000100022&script=sci_arttext
Rainwater H. 1963. Agricultural insect pest hitchhikers on aircraft. Proceedings of the Hawaiian Entomological Society 18: 303-310. (accessed 2015) http://scholarspace.manoa.hawaii.edu/bitstream/handle/10125/10881/18_303-310.pdf?sequence=1
Ratcliffe B and Paulsen P. 2008. The scarabaeoid beetles of Nebraska (Coleoptera: Scarabaeoidea). Bulletin of the University of Nebraska 22: 138-248.
Ratcliffe B, Cave R, Cano C. 2013. The Dynastinae scarab beetles of Mexico, Guatemala, and Belize (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska 27: 211-13.
Ratcliffe B, Jameson M, Smith A. 2002. Chapter 34. Scarabaeidae Latreille 1802, pp. 39-42. In, Arnett, Thomas, Skelley, and Frank (eds.), American Beetles. Polyphaga: Scarabaeoidea through Curculionoidea. Volume 2. CRC Press, New York.
Ratnasingham S and Hebert PDN. 2007. BOLD: The Barcode of Life Data System (www.barcodinglife.org). Molecular Ecology Notes 7:355–364. [DOI: 10.1111/j.1471- 8286.2006.01678.x]
Reding M and Klein M. 2007. Life history of oriental beetle and other scarabs, and occurrence of Tiphia vernalis in Ohio nurseries. Journal of Entomological Science 42 (3): 329-340.
Riley C and Howard L. 1893. An injurious Hawaiian beetle. Insect Life 6: 43.
Ritcher P. 1966. White grubs and their allies: a study of North American scarabaeoid larvae. Oregon State University Monographs, Studies in Entomology 4: 1-219.
Robinson M. 1948. A review of the species of Canthon inhabiting the United States (Scarabaeidae: Coleoptera). Transactions of the American Entomological Society 74: 83-100. (accessed 2015) http://www.jstor.org/stable/25077578
Roffey J. 1958. Observations on the biology of Trox procerus Har. (Coleoptera, Trogidae), a predator of eggs of the desert locust, Schistocerca gregaria (Forsk.). Bulletin of Entomological Research 49: 449-465.
Rowland J. 2003. Male horn dimorphism, phylogeny and systematics of rhinoceros beetles of the genus Xylotrupes (Scarabaeidae: Coleoptera). Australian Journal of Zoology: 51: 213-258.
Schreiner I and Nafus D. 1986. Accidental introductions of insect pests to Guam, 1945-1985. Proceedings of the Hawaiian Entomological Society 27: 45-52. (accessed 2015) http://scholarspace.manoa.hawaii.edu/bitstream/handle/10125/11207/27_45-52.pdf?sequence=1
Shetlar D and Niemczyk H. 1999. Black turfgrass Ataenius & Aphodius. Turfgrass Insect Fact Sheet Series, H.D.N. Books, Wooster, OH. (accessed 2015) http://entomology.osu.edu/schoolipm/IPMfiles/btafactsheet.pdf
Sikharulidze A. 1975. Biting pests of tung and their control. Subtropicheskie Kul'tury 5: 71-76.
Simmons L and Emlen J. 2008. No fecundity cost of female secondary sexual trait expression in the horned beetle Onthophagus sagittarius. Journal of Evolutionary Biology 21: 1227-1235. DOI: 10.1111/j.1420-9101.2008.01575.x
Simpson G. 1990. Immature stages of Protaetia fusca (Herbst). (Coleoptera: Scarabaeidae: Cetoniinae) with notes on its biology. Journal of the Australian Entomology Society 29: 67-73. (accessed 2015) http://onlinelibrary.wiley.com/doi/10.1111/j.1440-6055.1990.tb00317.x/pdf
Skelley P. 2012. The Asiatic garden beetle, Maladera castanea (Arrow, 1913) (Coleoptera; Scarabaeidae), a white grub pest new to Florida. Florida Department of Agriculture & Consumer Services, Division of Plant Industry Entomology Circular 425: 1-3. (accessed 2015) https://www.freshfromflorida.com/content/download/10805/141121/ent425.pdf
Skelley P. 2013. Asiatic garden beetle, Maladera castanea. University of Florida, Featured Creatures. (accessed 2015). http://entnemdept.ufl.edu/creatures/ORN/TURF/asiatic_garden_beetle.htm
Solis-Blanco A. 2002. The genus Canthon (Coleoptera: Scarabaeidae) in Costa Rica. Giornale Italiano di Entomologia 10: 1-68.
Stanaway M, Zalucki M, Gillespie P, Rodriguez C, Maynard G. 2001. Pest risk assessment of insects in sea cargo containers. Australian Journal of Entomology 40: 180-192.
Suehiro A, 1960. Insects and other arthropods from Midway Atoll. Hawaiian Entomological Society 17: 289-298. (accessed 2015) http://scholarspace.manoa.hawaii.edu/bitstream/handle/10125/10821/17_289-298.pdf
Suzuki M, Hor H, Tachibana M, Asano S. 1994. Bacillus thuringiensis strain Buibui for control of cupreous chafer, Anomala cuprea (Coleoptera: Scarabaeidae), in turfgrass and sweet potato. Biological Control 4: 361-365. DOI: 10.1006/bcon.1994.1045
Swezey O. 1920. Wireworms as cane pests in Hawaii. Hawaiian Planter's Record 23 (1): 1-4.
Thomas M. 1998. A flower beetle, Euphoria sepulcralis (Fabricius) (Insecta: Coleoptera: Scarabaeidae). Florida Department of Agriculture & Consumer Services, Division of Plant Industry Entomology Circular 386: 1-2. (accessed 2015) http://edis.ifas.ufl.edu/pdffiles/IN/IN75000.pdf
Toyama GM and Ikeda JK. 1976. An evaluation of fly predators at animal farms on leeward and central Oahu. Proceedings of the Hawaiian Entomological Society 22: 369-384.
Tyndale-Biscoe M. 1990. Common dung beetles in pastures of southeastern Australia. CSIRO Publishing, Clayton, Australia.
Tyndale-Biscoe M. 2006. The phenology of Onitis alexis (Coleoptera: Scarabaeidae) in the Araluen Valley: survival in a marginal environment. Australian Journal of Ecology 134: 431-443.
Van Dyke E. 1922. A study of the lucanid Coleoptera of the Hawaiian Islands. Proceedings of the Hawaiian Entomological Society 5: 39–49. (accessed 2015) http://archive.org/stream/proceedingsofhaw5192123hawa/proceedingsofhaw5192123hawa_djvu.txt
Venugopal K, Thomas S, Flemming A. 2012. Diversity and community structure of dung beetles (Coleoptera, Scarabaeinae) associated with semi-urban fragmented agricultural land in the Malabar coast in southern India. Journal of Threatened Taxa 4: 2685–2692. DOI: 10.11609/jott.o3074.2685-92
Visser D and Stals R. 2012. Temnorhynchus coronatus (Fabricius) (Scarabaeidae: Dynastinae), potentially a pest of sweet potato in South Africa. African Entomology 20: 402–407. DOI: 10.4001/003.020.0222
Vittum P, Villani M, Tashiro H. 1999. Turfgrass insects of the United States and Canada. Cornell University Press, Ithaca, NY, USA. 422 p.
Watschke T, Dernoeden P, Shetlar P. 1994. Managing Turfgrass Pests. Boca Raton, FL. CRC Press.
Weber P. 1954. Recent liberations of beneficial insects in Hawaii–IV. Proceedings of the Hawaiian Entomological Society 15: 635-638. (accessed 2015) http://hdl.handle.net/10125/14864
Weber P. 1955. Recent introductions for biological control in Hawaii—1. Proceedings of the Hawaiian Entomological Society 16: 162-164. (accessed 2015) http://hdl.handle.net/10125/14888
Williams X. 1931. Handbook of the Insects and Other Invertebrates of Hawaiian Sugarcane Fields. Honolulu, Hawaii: Hawaiian Sugar Planters' Association. Advertiser Publisher Co.
Wood L and Kaufman P. 2008. Euoniticellus intermedius (Coleoptera: Scarabaeidae: Scarabaeinae: tribe Coprini): its presence and relative abundance in cattle pastures in north-central Florida. Florida Entomologist 91: 128-130.(accessed 2015) http://journals.fcla.edu/flaent/article/view/75773/73431
Woodruff R and Beck B. 1989. The scarab beetles of Florida (Coleoptera: Scarabaeidae) part II. The May or June beetles (genus Phyllophaga). Arthropods of Florida and Neighboring Land Areas 13: 1-223.
Woodruff R. 1973. The scarab beetles of Florida (Coleoptera: Scarabaeidae) part I. The Laparosticti (Subfamilies: Scarabaeinae, Aphodiinae, Hybosorinae, Ochodaeinae, Geotrupinae, Acanthocerinae). Arthropods of Florida and Neighboring Land Areas 8: 1-220.
Woodruff R. 2006. The Asian mango flower beetle, Protaetia fusca (Herbst), and Euphoria sepulcralis (Fabricius) in Florida and the West Indies (Coleoptera: Scarabaeidae: Cetoniinae). Insecta Mundi 20: 227-232. (accessed 2015) http://digitalcommons.unl.edu/cgi/viewcontent.cgi?article=1113&context=insectamundi
Xu J, Liu Q, Jing H, Pang B, Yang J, Zhao G, Li H. 2002. Isolation of Escherichia coli O157:H7 from dung beetles Catharsius molossus. Microbial Immunology 47: 45-49.
Yokoyama T, Hasegawa M, Fujiie A, Sawada M, Noguchi K. 1998. Microbial control of scarab beetle larvae by a formulation of Metarhizium anisopliae (Deuteromycotina: Hyphomycetes) in a sweet potato field. Applied Entomology and Zoology 33: 215-218. (accessed 2015) http://ci.nii.ac.jp/naid/110001103219
Zidek J. 2013. Checklist and bibliography of the Trogidae (Coleoptera: Scarabaeoidea). Insecta Mundi 314: 1-38. full text (accessed 2015)